Viola pedunculata |
Viola lanceolata |
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California golden violet, johnny-jump-up, wild pansy, yellow pansy |
bog white violet, Howell's violet, lance-leaf or bog white violet, lance-leaf violet, strap-leaf violet, violette lancéolée, white bog violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–39 cm. | Plants perennial, acaulescent, stoloniferous, 5–30 cm; stolons pale, often rooting and leafy at nodes; rhizome slender, fleshy. |
Stems | 1–10+, decumbent, ascending, or erect, leafy proximally and distally, glabrous or puberulent, from shallow to deep-seated, enlarged rhizome with fleshy to subligneous roots. |
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Leaves | cauline; stipules ovate, linear-lanceolate, or oblanceolate, sometimes leaflike, margins entire or glandular-toothed, apex acute to acuminate; petiole 2.7–7.2 cm, usually finely puberulent, sometimes glabrate; blade deltate to ovate, 1–5.5 × 1–5.5 cm, base truncate, subcordate, or attenuate, margins crenate to serrate, ciliate, apex acute to obtuse, surfaces subglabrous or sparsely puberulent. |
basal, 2–6(–9), ascending to erect; stipules linear-lanceolate, margins entire or irregularly lacerate (at least distally), apex acute; petiole 2–12 cm, glabrous or pubescent; blade unlobed, lanceolate or narrowly elliptic to nearly linear, 2.5–12 × 0.7–2.5 cm, longer than wide, base attenuate, margins serrate, mostly eciliate, apex acute, mucronulate, surfaces glabrous. |
Peduncles | 2.9–20 cm, sparsely to densely puberulent. |
2–17 cm, usually pubescent. |
Flowers | sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–3 mm; petals golden yellow adaxially, upper 2 reddish brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 10–20 mm, spur dark reddish brown, gibbous, 2–4 mm; style head bearded; cleistogamous flowers absent. |
sepals ovate to lanceolate, margins mostly eciliate, auricles 1–2 mm; petals white on both surfaces, lowest and sometimes lateral 2 purple-veined, lateral 2 sparsely bearded or beardless, lowest 7–12 mm, spur white, gibbous, 1–2 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
Capsules | ellipsoid, 5–11 mm, glabrous. |
ellipsoid, 5–8 mm, glabrous. |
Seeds | dark brown or black, shiny, 2.7 mm. |
beige to bronze, 1.5–2.5 mm. |
2n | = 12. |
= 24. |
Viola pedunculata |
Viola lanceolata |
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Phenology | Flowering Feb–Apr. | Flowering Mar–May. |
Habitat | Open, grassy coastal and inland slopes and hillsides, usually in full sun, chaparral, foothill and oak woodland | Open to semi-open wet areas, bogs, meadows, pond and lake shores, stream banks, seasonally inundated depressions |
Elevation | 0–1000 m (0–3300 ft) | 0–2000 m (0–6600 ft) |
Distribution |
CA; Mexico (Baja California)
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AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; OR; PA; RI; SC; TN; TX; VA; VT; WA; WI; WV; BC; NB; NF; NS; ON; PE; QC; SPM
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Discussion | The stems of Viola pedunculata arise from an enlarged, subterranean, spongy or fibrous rhizome. Often, these rhizome structures are deep seated; it is unknown how they get so deeply buried. The anther appendages of V. pedunculata are hairy distally, a characteristic not known to occur in other members of the V. purpurea complex. Larvae of the federally listed Callippe silverspot butterfly [Speyeria callippe (Boisduval) callippe] feed only on Viola pedunculata. Plants with leaves reported to be smaller, thinner, deltate, mostly longer than wide, with yellow petals (versus orange for Viola pedunculata var. pedunculata), style 2.1 mm (versus 2.9 mm for var. pedunculata), from the Pinnacles region in San Benito County, California, have been called subsp. tenuifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola lanceolata occurs in small colonies; individual plants are interconnected by stolons. Viola lanceolata subsp. vittata was recognized by N. H. Russell (1955) based on its linear leaf blade shape. The range in leaf blade shape appears to have no distinct line of demarcation. Some believe that leaf shape differences and near restriction to the coastal plain support recognition at some level. Studies are necessary to resolve this issue. J. H. Shultze (1946) reported that Viola lanceolata was introduced into Washington in the early 1900s primarily from Cape Cod and Wisconsin as a result of importation of cranberry vines. In British Columbia, it is known from Lulu Island, where it was introduced from eastern North America (G. W. Douglas et al. 1998–2002, vol. 5). Viola lanceolata reportedly hybridizes with V. primulifolia var. primulifolia (= V. ×modesta House) and V. macloskeyi (= V. ×sublanceolata House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 144. | FNA vol. 6, p. 135. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. pedunculata subsp. tenuifolia | V. lanceolata subsp. vittata, V. lanceolata var. vittata, V. vittata |
Name authority | Torrey & A. Gray: Fl. N. Amer. 1: 141. (1838) | Linnaeus: Sp. Pl. 2: 934. (1753) |
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