Viola pedunculata |
Viola douglasii |
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California golden violet, johnny-jump-up, wild pansy, yellow pansy |
Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–39 cm. | Plants perennial, caulescent, not stoloniferous, 3–20 cm. |
Stems | 1–10+, decumbent, ascending, or erect, leafy proximally and distally, glabrous or puberulent, from shallow to deep-seated, enlarged rhizome with fleshy to subligneous roots. |
1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex. |
Leaves | cauline; stipules ovate, linear-lanceolate, or oblanceolate, sometimes leaflike, margins entire or glandular-toothed, apex acute to acuminate; petiole 2.7–7.2 cm, usually finely puberulent, sometimes glabrate; blade deltate to ovate, 1–5.5 × 1–5.5 cm, base truncate, subcordate, or attenuate, margins crenate to serrate, ciliate, apex acute to obtuse, surfaces subglabrous or sparsely puberulent. |
basal and cauline; basal: 1–6, bipinnately compound, leaflets 3–5; stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate; petiole 5–6.8 cm, glabrous or puberulent; blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute; petiole 0.9–4 cm; blade 1.1–4.1 × 1–3.6 cm. |
Peduncles | 2.9–20 cm, sparsely to densely puberulent. |
2–12.5 cm, glabrous or puberulent. |
Flowers | sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–3 mm; petals golden yellow adaxially, upper 2 reddish brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 10–20 mm, spur dark reddish brown, gibbous, 2–4 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm; style head bearded; cleistogamous flowers absent. |
Capsules | ellipsoid, 5–11 mm, glabrous. |
spherical to oblong, 5–12 mm, glabrous. |
Seeds | dark brown or black, shiny, 2.7 mm. |
light brown, 2.8–3.3 mm. |
2n | = 12. |
= 24, 48. |
Viola pedunculata |
Viola douglasii |
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Phenology | Flowering Feb–Apr. | Flowering Feb–Jul. |
Habitat | Open, grassy coastal and inland slopes and hillsides, usually in full sun, chaparral, foothill and oak woodland | Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) |
Elevation | 0–1000 m (0–3300 ft) | 20–2300 m (100–7500 ft) |
Distribution |
CA; Mexico (Baja California)
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CA; OR; Mexico (Baja California)
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Discussion | The stems of Viola pedunculata arise from an enlarged, subterranean, spongy or fibrous rhizome. Often, these rhizome structures are deep seated; it is unknown how they get so deeply buried. The anther appendages of V. pedunculata are hairy distally, a characteristic not known to occur in other members of the V. purpurea complex. Larvae of the federally listed Callippe silverspot butterfly [Speyeria callippe (Boisduval) callippe] feed only on Viola pedunculata. Plants with leaves reported to be smaller, thinner, deltate, mostly longer than wide, with yellow petals (versus orange for Viola pedunculata var. pedunculata), style 2.1 mm (versus 2.9 mm for var. pedunculata), from the Pinnacles region in San Benito County, California, have been called subsp. tenuifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 144. | FNA vol. 6, p. 129. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. pedunculata subsp. tenuifolia | V. chrysantha |
Name authority | Torrey & A. Gray: Fl. N. Amer. 1: 141. (1838) | Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) |
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