Viola pedunculata |
Viola canina |
|
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California golden violet, johnny-jump-up, wild pansy, yellow pansy |
dog violet, heath dog-violet |
|
Habit | Plants perennial, caulescent, not stoloniferous, 5–39 cm. | Plants perennial, caulescent, not stoloniferous, 2–40 cm. |
Stems | 1–10+, decumbent, ascending, or erect, leafy proximally and distally, glabrous or puberulent, from shallow to deep-seated, enlarged rhizome with fleshy to subligneous roots. |
1–5, ascending to erect, glabrous, on caudex from subligneous rhizome. |
Leaves | cauline; stipules ovate, linear-lanceolate, or oblanceolate, sometimes leaflike, margins entire or glandular-toothed, apex acute to acuminate; petiole 2.7–7.2 cm, usually finely puberulent, sometimes glabrate; blade deltate to ovate, 1–5.5 × 1–5.5 cm, base truncate, subcordate, or attenuate, margins crenate to serrate, ciliate, apex acute to obtuse, surfaces subglabrous or sparsely puberulent. |
cauline; stipules linear to lanceolate, margins ± lacerate to subentire, points often gland-tipped, apex acute; petiole 0.3–2.5 cm, glabrous; blade ovate to narrowly ovate, 1.2–5.2 × 1–2.7 cm, base cordate or deeply cordate to truncate, margins crenate, eciliate, apex acute to obtuse, abaxial surface glabrous or sparsely pubescent on veins, adaxial surface glabrous or sparsely pubescent. |
Peduncles | 2.9–20 cm, sparsely to densely puberulent. |
1–10 cm, glabrous or sparsely puberulent. |
Flowers | sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–3 mm; petals golden yellow adaxially, upper 2 reddish brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 10–20 mm, spur dark reddish brown, gibbous, 2–4 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate, margins eciliate, auricles 2–3 mm; petals usually blue- to gray-violet on both surfaces, rarely white, white basally, lower 3 dark violet-veined, lateral 2 bearded, lowest 15–25 mm, spur light yellow to light green or white, gibbous to usually elongated, 3–5 mm; style head beardless; cleistogamous flowers axillary. |
Capsules | ellipsoid, 5–11 mm, glabrous. |
ovoid, 7.5–9 mm, glabrous. |
Seeds | dark brown or black, shiny, 2.7 mm. |
light to dark brown; 1.7–2.1 mm. |
2n | = 12. |
= 40. |
Viola pedunculata |
Viola canina |
|
Phenology | Flowering Feb–Apr. | Flowering May–Jul. |
Habitat | Open, grassy coastal and inland slopes and hillsides, usually in full sun, chaparral, foothill and oak woodland | Sunny to shady, dry to slightly moist ground, heath lands, grazed or mown grassland, dunes, rock ledges, open woodland, disturbed areas (roadsides, railway banks, gravel pits), littoral vegetation of lakes and streams |
Elevation | 0–1000 m (0–3300 ft) | 20–300 m (100–1000 ft) |
Distribution |
CA; Mexico (Baja California)
|
Greenland; Europe; Asia |
Discussion | The stems of Viola pedunculata arise from an enlarged, subterranean, spongy or fibrous rhizome. Often, these rhizome structures are deep seated; it is unknown how they get so deeply buried. The anther appendages of V. pedunculata are hairy distally, a characteristic not known to occur in other members of the V. purpurea complex. Larvae of the federally listed Callippe silverspot butterfly [Speyeria callippe (Boisduval) callippe] feed only on Viola pedunculata. Plants with leaves reported to be smaller, thinner, deltate, mostly longer than wide, with yellow petals (versus orange for Viola pedunculata var. pedunculata), style 2.1 mm (versus 2.9 mm for var. pedunculata), from the Pinnacles region in San Benito County, California, have been called subsp. tenuifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola canina was possibly introduced in Greenland (T. Marcussen and T. Karlsson et al. 2010). Hybrids of Viola canina are known with the European species V. mirabilis Linnaeus, V. pumila Chaix, V. reichenbachiana Jordan ex Boreau, V. riviniana Reichenbach, V. rupestris F. W. Schmidt (both subspecies), V. stagnina Kitaibel, and V. uliginosa Besser. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 144. | FNA vol. 6, p. 127. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. pedunculata subsp. tenuifolia | V. canina var. montana, V. montana, V. nemoralis |
Name authority | Torrey & A. Gray: Fl. N. Amer. 1: 141. (1838) | Linnaeus: Sp. Pl. 2: 935. (1753) |
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