Viola pedunculata
Viola beckwithii
California golden violet, johnny-jump-up, wild pansy, yellow pansy
Beckwith's or Great Basin violet, Beckwith's violet, Great Basin violet, sagebrush pansy, sagebrush violet
1–10+, decumbent, ascending, or erect, leafy proximally and distally, glabrous or puberulent, from shallow to deep-seated, enlarged rhizome with fleshy to subligneous roots.
1–3, decumbent, ascending, or erect, ca. 1/2 subterranean, glabrous or usually puberulent, on single, short, vertical, deep-seated caudex.
cauline;
stipules ovate, linear-lanceolate, or oblanceolate, sometimes leaflike, margins entire or glandular-toothed, apex acute to acuminate;
petiole 2.7–7.2 cm, usually finely puberulent, sometimes glabrate;
blade deltate to ovate, 1–5.5 × 1–5.5 cm, base truncate, subcordate, or attenuate, margins crenate to serrate, ciliate, apex acute to obtuse, surfaces subglabrous or sparsely puberulent.
basal and cauline;
basal: 1–6 per caudex, palmately compound, ± 2-ternate or 3-ternate, leaflets 3;
stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute;
petiole 2–10.5 cm, usually puberulent;
blade ovate to deltate, 2.4–5 × 3.5–4.5 cm, base tapered, ultimate leaflets dissected into oblong, elliptic, lanceolate, or oblanceolate lobes 1–7 mm wide, lobe margins entire, usually ciliate, apex acute to obtuse, mucronulate, surfaces usually puberulent, seldom glabrous, abaxial surface without prominent vein parallel to each margin;
cauline similar to basal except: stipules linear, apex acuminate;
petiole 2–5.7 cm, usually puberulent, rarely glabrous;
blade 1–2.7 × 1.5–3 cm.
2.9–20 cm, sparsely to densely puberulent.
1.5–10.6(–15.7) cm, usually puberulent, seldom glabrous.
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–3 mm;
petals golden yellow adaxially, upper 2 reddish brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 10–20 mm, spur dark reddish brown, gibbous, 2–4 mm;
style head bearded; cleistogamous flowers absent.
sepals lanceolate, margins eciliate, auricles 0.1–1 mm;
petals dark reddish violet on both surfaces, lower 3 usually lilac, rarely white or whitish, lateral 2 bearded, with yellow patch basally, dark reddish violet-veined, lowest 10–22 mm, with yellow patch, dark reddish violet-veined, spur whitish or yellowish, tinged purple, gibbous, 0.5–2 mm;
style head bearded; cleistogamous flowers absent.
ellipsoid, 5–11 mm, glabrous.
oblong-ovoid, 7–12 mm, glabrous.
dark brown or black, shiny, 2.7 mm.
brown, 3–4 mm.
= 12.
= 24.
Viola pedunculata
Viola beckwithii
The stems of Viola pedunculata arise from an enlarged, subterranean, spongy or fibrous rhizome. Often, these rhizome structures are deep seated; it is unknown how they get so deeply buried. The anther appendages of V. pedunculata are hairy distally, a characteristic not known to occur in other members of the V. purpurea complex.
Larvae of the federally listed Callippe silverspot butterfly [Speyeria callippe (Boisduval) callippe] feed only on Viola pedunculata.
Plants with leaves reported to be smaller, thinner, deltate, mostly longer than wide, with yellow petals (versus orange for Viola pedunculata var. pedunculata), style 2.1 mm (versus 2.9 mm for var. pedunculata), from the Pinnacles region in San Benito County, California, have been called subsp. tenuifolia.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
In some populations of Viola beckwithii, the three lower petals are white with a yellow area proximally (V. B. Baird 1942). Leaves have been described as palmately biternate or triternate (L. Abrams and R. S. Ferris 1923–1960, vol. 3), ternately decompound into linear segments (C. L. Hitchcock et al. 1955–1969, vol. 3), palmately three-parted then bipinnately parted into ultimate linear or spatulate segments (P. A. Munz 1959), and palmately about three times three-parted into linear or spatulate-linear segments (W. L. Jepson 1951). Some populations in northern California are nearly or completely glabrous, which M. S. Baker recognized as var. glabrata.
Viola beckwithii is reported to hybridize with V. utahensis (G. Davidse 1976). Observed pollinators of V. beckwithii in Utah include Apis mellifera Linnaeus and Anthophora ursina Cresson (Davidse).
When Cottam described Viola bonnevillensis, he suggested that it could be a hybrid between V. beckwithii and V. utahensis, and G. Davidse (1976) concurred. The type specimen of V. bonnevillensis (Cottam 7067, UT) was examined by R. J. Little. Because no similar forms are known to have been collected since 1939, it is presumed that this taxon is a hybrid.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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