Viola pedunculata |
Viola bakeri |
|
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California golden violet, johnny-jump-up, wild pansy, yellow pansy |
Baker's violet, yellow prairie violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–39 cm. | Plants perennial, caulescent, not stoloniferous, 3–30 cm. |
Stems | 1–10+, decumbent, ascending, or erect, leafy proximally and distally, glabrous or puberulent, from shallow to deep-seated, enlarged rhizome with fleshy to subligneous roots. |
1–4, usually erect, sometimes prostrate or decumbent, leafy proximally and distally, usually puberulent, from usually vertical, subligneous rhizome. |
Leaves | cauline; stipules ovate, linear-lanceolate, or oblanceolate, sometimes leaflike, margins entire or glandular-toothed, apex acute to acuminate; petiole 2.7–7.2 cm, usually finely puberulent, sometimes glabrate; blade deltate to ovate, 1–5.5 × 1–5.5 cm, base truncate, subcordate, or attenuate, margins crenate to serrate, ciliate, apex acute to obtuse, surfaces subglabrous or sparsely puberulent. |
basal and cauline; basal: 1–4; stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, acute to acuminate; petiole 1–15.4 cm, glabrous or puberulent; blade lanceolate, oblanceolate, or elliptic, rarely ovate, 1.8–8.8 × 0.7–3.9 cm, thin, base attenuate, often oblique, margins usually entire, sometimes with a few sharp teeth or crenulate, ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent on margins or veins; cauline similar to basal except: stipules ovate to lanceolate, margins entire or lacerate, sometimes with glandular projections, apex with 2–3 projections; petiole 1.5–7.5 cm; blade 1.9–6.7 × 0.5–1.6 cm. |
Peduncles | 2.9–20 cm, sparsely to densely puberulent. |
1.5–11.6 cm, glabrous or pubescent. |
Flowers | sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–3 mm; petals golden yellow adaxially, upper 2 reddish brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 10–20 mm, spur dark reddish brown, gibbous, 2–4 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 6–14 mm, spur yellow, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary. |
Capsules | ellipsoid, 5–11 mm, glabrous. |
spherical to ovoid, 5–10 mm, usually glabrous, rarely finely puberulent. |
Seeds | dark brown or black, shiny, 2.7 mm. |
light to medium brown or dark red-brown, 2.6–3.1 mm, elaiosome not covering funiculus. |
2n | = 12. |
= 48. |
Viola pedunculata |
Viola bakeri |
|
Phenology | Flowering Feb–Apr. | Flowering May–Jul. |
Habitat | Open, grassy coastal and inland slopes and hillsides, usually in full sun, chaparral, foothill and oak woodland | Wet and dry places in openings of coniferous forests |
Elevation | 0–1000 m (0–3300 ft) | 900–3800 m (3000–12500 ft) |
Distribution |
CA; Mexico (Baja California)
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CA; NV; OR; WA
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Discussion | The stems of Viola pedunculata arise from an enlarged, subterranean, spongy or fibrous rhizome. Often, these rhizome structures are deep seated; it is unknown how they get so deeply buried. The anther appendages of V. pedunculata are hairy distally, a characteristic not known to occur in other members of the V. purpurea complex. Larvae of the federally listed Callippe silverspot butterfly [Speyeria callippe (Boisduval) callippe] feed only on Viola pedunculata. Plants with leaves reported to be smaller, thinner, deltate, mostly longer than wide, with yellow petals (versus orange for Viola pedunculata var. pedunculata), style 2.1 mm (versus 2.9 mm for var. pedunculata), from the Pinnacles region in San Benito County, California, have been called subsp. tenuifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
M. S. Baker (1957) wrote that Viola bakeri flowers were often without a brownish tinge on the back. Subsequent collections have shown that the upper two petals of V. bakeri are often brownish purple abaxially and thus this characteristic cannot be used to distinguish V. bakeri from V. praemorsa, as previously done (P. A. Munz 1959; C. L. Hitchcock et al. 1955–1969, vol. 3). Greene wrote in his description of Viola bakeri that the whole plant was glabrous; he did not mention leaf margins. M. S. Baker (1957) examined the type specimen at UC and apparently was the first to document that its leaf margins are entire. He also noted that under magnification the leaves are ciliate and more or less puberulent. D. M. Fabijan et al. (1987) stated that margins were always entire and ciliate and more or less pubescent throughout. Some collections of V. bakeri from California have leaves with a sharp point or two on the margin, or crenulations on some portion of the basal or cauline blade margins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 144. | FNA vol. 6, p. 121. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. pedunculata subsp. tenuifolia | V. bakeri subsp. shastensis, V. nuttallii var. bakeri |
Name authority | Torrey & A. Gray: Fl. N. Amer. 1: 141. (1838) | Greene: Pittonia 3: 307. (1898) |
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