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California golden violet, johnny-jump-up, wild pansy, yellow pansy

Le Conte's violet, sand violet, violette affine

Habit Plants perennial, caulescent, not stoloniferous, 5–39 cm. Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age.
Stems

1–10+, decumbent, ascending, or erect, leafy proximally and distally, glabrous or puberulent, from shallow to deep-seated, enlarged rhizome with fleshy to subligneous roots.

Leaves

cauline;

stipules ovate, linear-lanceolate, or oblanceolate, sometimes leaflike, margins entire or glandular-toothed, apex acute to acuminate;

petiole 2.7–7.2 cm, usually finely puberulent, sometimes glabrate;

blade deltate to ovate, 1–5.5 × 1–5.5 cm, base truncate, subcordate, or attenuate, margins crenate to serrate, ciliate, apex acute to obtuse, surfaces subglabrous or sparsely puberulent.

basal, 1–6, ascending to erect;

stipules lanceolate, margins entire or fimbriate, apex acute;

petiole 2–10 cm, glabrous;

blade green abaxially, unlobed, narrowly to broadly ovate or narrowly deltate, 1.5–10 × 1.5–10 cm, not fleshy, base cordate to broadly cordate or almost truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse, surfaces sparsely pubescent adaxially, rarely glabrous.

Peduncles

2.9–20 cm, sparsely to densely puberulent.

3–15 cm, glabrous or pubescent.

Flowers

sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–3 mm;

petals golden yellow adaxially, upper 2 reddish brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 10–20 mm, spur dark reddish brown, gibbous, 2–4 mm;

style head bearded; cleistogamous flowers absent.

sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm;

petals lavender-violet to dull reddish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, lowest 10–22 mm, usually obviously bearded, rarely beardless, spur white or same color as petals, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers from prostrate to ascending peduncles.

Capsules

ellipsoid, 5–11 mm, glabrous.

often reddish or purplish-flecked or green, ellipsoid, 5–10 mm, glabrous or puberulent.

Seeds

dark brown or black, shiny, 2.7 mm.

beige, mottled to bronze, 1.5–2.5 mm.

2n

= 12.

= 54.

Viola pedunculata

Viola affinis

Phenology Flowering Feb–Apr. Flowering Apr–Jun.
Habitat Open, grassy coastal and inland slopes and hillsides, usually in full sun, chaparral, foothill and oak woodland Open or wooded wet areas, meadows, stream banks, thickets, shores of lakes, seasonally dry areas.
Elevation 0–1000 m (0–3300 ft) 100–2000 m (300–6600 ft)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; DC; DE; IL; IN; MA; MD; MI; MN; NH; NJ; NY; OH; PA; RI; VA; VT; WI; WV; ON; QC
[BONAP county map]
Discussion

The stems of Viola pedunculata arise from an enlarged, subterranean, spongy or fibrous rhizome. Often, these rhizome structures are deep seated; it is unknown how they get so deeply buried. The anther appendages of V. pedunculata are hairy distally, a characteristic not known to occur in other members of the V. purpurea complex.

Larvae of the federally listed Callippe silverspot butterfly [Speyeria callippe (Boisduval) callippe] feed only on Viola pedunculata.

Plants with leaves reported to be smaller, thinner, deltate, mostly longer than wide, with yellow petals (versus orange for Viola pedunculata var. pedunculata), style 2.1 mm (versus 2.9 mm for var. pedunculata), from the Pinnacles region in San Benito County, California, have been called subsp. tenuifolia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

L. E. McKinney (1992) considered Viola affinis, and much of what botanists had called V. nephrophylla, to be essentially the same taxon. After studying additional specimens, reviewing literature (H. E. Ballard 1994; A. Haines 2001b), and discussions with others (J. Cayouette, H. E. Ballard, A. Haines, pers. comm.), he chose to maintain these as separate taxa. Reports of V. affinis in the Gulf coastal states based on specimens or photographs are usually attributable to V. missouriensis.

Viola affinis reportedly hybridizes with V. hirsutula (= V. ×consobrina House), V. cucullata (= V. ×consocia House), V. brittoniana (= V. ×davisii House), V. sororia (= V. ×filicetorum Greene [as species]), V. sagittata var. sagittata (= V. ×hollickii House), and V. nephrophylla (= V. ×subaffinis House).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 144. FNA vol. 6, p. 120.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. pedunculata subsp. tenuifolia V. affinis var. subarctica, V. crenulata, V. sororia subsp. affinis, V. sororia var. affinis, V. subviscosa, V. venustula
Name authority Torrey & A. Gray: Fl. N. Amer. 1: 141. (1838) Leconte: Ann. Lyceum Nat. Hist. New York 2: 138. (1826)
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