Viola pedunculata |
Viola adunca |
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California golden violet, johnny-jump-up, wild pansy, yellow pansy |
Cascades early blue violet, common periwinkle, early blue violet, hook-spur violet, hookedspur violet, large periwinkle, sand violet, western dog violet, wild dog violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–39 cm. | Plants perennial, caulescent, not stoloniferous, 1.8–30(–35) cm. | ||||
Stems | 1–10+, decumbent, ascending, or erect, leafy proximally and distally, glabrous or puberulent, from shallow to deep-seated, enlarged rhizome with fleshy to subligneous roots. |
1–5, erect, ascending, or decumbent, sometimes later reclining to nearly prostrate, glabrous or puberulent, on caudex from subligneous rhizome. |
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Leaves | cauline; stipules ovate, linear-lanceolate, or oblanceolate, sometimes leaflike, margins entire or glandular-toothed, apex acute to acuminate; petiole 2.7–7.2 cm, usually finely puberulent, sometimes glabrate; blade deltate to ovate, 1–5.5 × 1–5.5 cm, base truncate, subcordate, or attenuate, margins crenate to serrate, ciliate, apex acute to obtuse, surfaces subglabrous or sparsely puberulent. |
basal and cauline; basal: 1–4; stipules linear to linear-lanceolate, margins entire or laciniate with gland-tipped projections, apex acute to acuminate; petiole 0.5–13.5 cm, glabrous or puberulent; blade usually ovate or ovate-deltate to ovate-orbiculate, sometimes ± reniform or oblong, 0.5–6.9 × 0.4–5 cm, base cordate, subcordate, truncate, or attenuate, usually decurrent on petiole, margins crenate to crenulate or entire, ciliate or eciliate, apex acute to obtuse, surfaces glabrous or sparsely to densely puberulent; cauline similar to basal except: stipule margins lacerate to laciniate; petiole 0.5–6.5 cm; blade 0.6–5.5 × 0.4–4.7 cm. |
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Peduncles | 2.9–20 cm, sparsely to densely puberulent. |
1.7–13.8 cm, glabrous or puberulent. |
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Flowers | sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–3 mm; petals golden yellow adaxially, upper 2 reddish brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 10–20 mm, spur dark reddish brown, gibbous, 2–4 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate, margins ciliate or eciliate, auricles not enlarged in fruit, 0.5–2 mm; petals light- to deep- to lavender-violet on both surfaces, rarely white, lower 3 usually white basally, dark violet-veined, lateral 2 (and sometimes upper 2) bearded, lowest 7–17(–23) mm, spur purple to violet or white, elongated, 5–7 mm, tip straight or pointed, curved up or lateral; style head sparsely to densely bearded, sometimes beardless; cleistogamous flowers axillary. |
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Capsules | ellipsoid, 5–11 mm, glabrous. |
short-ovoid, 6–11 mm, glabrous. |
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Seeds | dark brown or black, shiny, 2.7 mm. |
dark brown to olive-black, 1.5–2 mm. |
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2n | = 12. |
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Viola pedunculata |
Viola adunca |
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Phenology | Flowering Feb–Apr. | |||||
Habitat | Open, grassy coastal and inland slopes and hillsides, usually in full sun, chaparral, foothill and oak woodland | |||||
Elevation | 0–1000 m (0–3300 ft) | |||||
Distribution |
CA; Mexico (Baja California)
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AK; AZ; CA; CO; IA; ID; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OR; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NS; NT; NU; ON; QC; SK; YT
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Discussion | The stems of Viola pedunculata arise from an enlarged, subterranean, spongy or fibrous rhizome. Often, these rhizome structures are deep seated; it is unknown how they get so deeply buried. The anther appendages of V. pedunculata are hairy distally, a characteristic not known to occur in other members of the V. purpurea complex. Larvae of the federally listed Callippe silverspot butterfly [Speyeria callippe (Boisduval) callippe] feed only on Viola pedunculata. Plants with leaves reported to be smaller, thinner, deltate, mostly longer than wide, with yellow petals (versus orange for Viola pedunculata var. pedunculata), style 2.1 mm (versus 2.9 mm for var. pedunculata), from the Pinnacles region in San Benito County, California, have been called subsp. tenuifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Viola adunca is polymorphic with over 50 named taxa. G. D. McPherson and J. G. Packer (1974) reported that diploid, triploid, and tetraploid races of V. adunca occur in Canada and northwestern United States. They found that diploid and tetraploid chromosome races can be distinguished morphologically based on style beards and on the size of guard cells and pollen grains and recommended taxonomic recognition of both races. In diploid races, the style projections are more or less cylindrical and about one-sixth the width of the style; in tetraploid races they are short-conical or globular, and about one-tenth or less than the width of the style beard. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 144. | FNA vol. 6, p. 119. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. pedunculata subsp. tenuifolia | Lophion aduncum, V. canina var. adunca | ||||
Name authority | Torrey & A. Gray: Fl. N. Amer. 1: 141. (1838) | Smith: in A. Rees, Cycl. 37: Viola no. 63 | ||||
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