Viola pedatifida |
Viola nephrophylla |
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crow-foot violet, prairie or crowfoot or larkspur violet, prairie violet, violette pédatifide |
kidney-leaf violet, Leconte violet, northern bog violet, violette néphrophylle |
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Habit | Plants perennial, acaulescent, not stoloniferous, 5–30 cm; rhizome thick, fleshy. | Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age. |
Leaves | basal, 2–11, ascending to erect, 5–9-lobed; stipules linear-lanceolate, margins entire, apex acute; petiole 3–16 cm, pubescent; blade similar in width and shape, lobes lanceolate, spatulate, falcate, or linear, 1–7 × 2–8 cm, base truncate to reniform, margins entire, ciliate, apex acute to obtuse, mucronulate, surfaces pubescent, hairs sometimes concentrated on veins. |
basal, 4–7, ascending to erect; stipules lanceolate, margins entire or fimbriate, apex acute; petiole 2–25 cm, usually glabrous; blade usually grayish green or purplish green abaxially, unlobed, ovate, reniform, or broadly reniform to orbiculate, 1–7 × 1–7 cm, somewhat fleshy, base broadly cordate or reniform to ± truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually glabrous, rarely sparsely pubescent. |
Peduncles | 5–18 cm, glabrous or pubescent. |
3–25 cm, usually glabrous. |
Flowers | sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to soft reddish violet on both surfaces, lower 3 white basally, dark violet-veined, lateral 2 and lowest usually bearded, lowest 10–25 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on ascending to erect peduncles. |
sepals ovate, margins usually eciliate, auricles 1–2 mm; petals deep bluish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, upper 2 sometimes sparsely bearded, lowest densely bearded or beardless, 10–28 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on erect to ascending peduncles. |
Capsules | ellipsoid, 10–15 mm, glabrous. |
ovoid, 5–10 mm, glabrous. |
Seeds | beige, mottled to bronze, 1.5–2.5 mm. |
beige to brown or dark brown, 1.5–2.5 mm. |
2n | = 54. |
= 54. |
Viola pedatifida |
Viola nephrophylla |
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Phenology | Flowering Mar–Jun. | Flowering Jan–Sep. |
Habitat | Prairies, grasslands, disturbed ground, dry gravelly hills | Wet habitats in saturated soil in prairies, pastures, bogs, fens, sedge meadows, rocky shores of lakes and streams, limestone outcrops, gravelly calcareous stream beds |
Elevation | 500–1000 m (1600–3300 ft) | 100–3000 m (300–9800 ft) |
Distribution |
AR; AZ; CO; IA; IL; IN; KS; MI; MN; MO; MT; ND; NE; NM; OH; OK; SD; VA; WI; WY; AB; MB; ON; SK
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AR; AZ; CA; CO; CT; IA; ID; IL; KS; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OH; OR; PA; SD; TX; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT
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Discussion | Viola pedatifida was reported historically from, and recently rediscovered in, the Appalachian shale barrens of Virginia (T. Wieboldt, pers. comm.). Viola pedatifida reportedly hybridizes with V. sororia (= V. ×bernardii Greene). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola nephrophylla has the widest distribution in North America of any of the acaulescent blue violets. There is some question whether the species we now know as Viola nephrophylla fits with the protologue and type specimen(s). McKinney examined the type designated by Greene; it appears that what most botanists have called V. nephrophylla may be an undescribed taxon. N. L. Gil-Ad (1997) believed Greene’s type contained plants that were genetically impure based on seed coat micromorphology and suggested the specimens might be a mixture of both orthospecies and introgressants. Viola mccabeiana was described by M. S. Baker in 1940 from collections made in British Columbia. Later, Baker (1949b) discussed the reasons why the name V. mccabeiana needed to be abandoned. C. L. Hitchcock et al. (1955–1969, vol. 3) treated V. mccabeiana as synonymous with V. nephrophylla var. cognata. While Viola mccabeiana is recognized in Canada (L. Brouillet et al., http://data.canadensys.net/vascan/), we believe it is best included within V. nephrophylla. Viola nephrophylla reportedly hybridizes with V. cucullata (= V. ×insessa House), V. sororia var. sororia (= V. ×napae House), and V. affinis (= V. ×subaffinis House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 144. | FNA vol. 6, p. 139. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. delphiniifolia, V. palmata var. pedatifida | V. arizonica, V. austiniae, V. cognata, V. crassula, V. galacifolia, V. lunellii, V. mccabeiana, V. nephrophylla var. arizonica, V. nephrophylla var. cognata, V. peramoena, V. pratincola, V. prionosepala, V. retusa, V. subjuncta, V. vagula |
Name authority | G. Don: Gen. Hist. 1: 320. (1831) | Greene: Pittonia 3: 144. (1896) |
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