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crow-foot violet, prairie or crowfoot or larkspur violet, prairie violet, violette pédatifide

Howell's violet, violette de Howell

Habit Plants perennial, acaulescent, not stoloniferous, 5–30 cm; rhizome thick, fleshy. Plants perennial, caulescent, not stoloniferous, 2–44 cm.
Stems

1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome.

Leaves

basal, 2–11, ascending to erect, 5–9-lobed;

stipules linear-lanceolate, margins entire, apex acute;

petiole 3–16 cm, pubescent;

blade similar in width and shape, lobes lanceolate, spatulate, falcate, or linear, 1–7 × 2–8 cm, base truncate to reniform, margins entire, ciliate, apex acute to obtuse, mucronulate, surfaces pubescent, hairs sometimes concentrated on veins.

basal and cauline;

basal: 1–6;

stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate;

petiole 4–15 cm, glabrous or sparsely puberulent;

blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent;

cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate;

petiole 1–4.5 cm;

blade 2–5.1 × 1.2–5.8 cm.

Peduncles

5–18 cm, glabrous or pubescent.

2.8–17.8 cm, glabrous or sparsely pubescent.

Flowers

sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm;

petals light to soft reddish violet on both surfaces, lower 3 white basally, dark violet-veined, lateral 2 and lowest usually bearded, lowest 10–25 mm, spur same color as petals, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers on ascending to erect peduncles.

sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm;

petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight;

style head bearded; cleistogamous flowers axillary.

Capsules

ellipsoid, 10–15 mm, glabrous.

ellipsoid, 7–11 mm, glabrous.

Seeds

beige, mottled to bronze, 1.5–2.5 mm.

light brown, 2.1–2.8 mm.

2n

= 54.

= 40, 80.

Viola pedatifida

Viola howellii

Phenology Flowering Mar–Jun. Flowering Apr–Jul.
Habitat Prairies, grasslands, disturbed ground, dry gravelly hills Moist, shady areas, coniferous forests, prairies, along streams
Elevation 500–1000 m (1600–3300 ft) 50–1500 m (200–4900 ft)
Distribution
from FNA
AR; AZ; CO; IA; IL; IN; KS; MI; MN; MO; MT; ND; NE; NM; OH; OK; SD; VA; WI; WY; AB; MB; ON; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola pedatifida was reported historically from, and recently rediscovered in, the Appalachian shale barrens of Virginia (T. Wieboldt, pers. comm.).

Viola pedatifida reportedly hybridizes with V. sororia (= V. ×bernardii Greene).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 144. FNA vol. 6, p. 134.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. delphiniifolia, V. palmata var. pedatifida
Name authority G. Don: Gen. Hist. 1: 320. (1831) A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887)
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