Viola pedatifida
Viola biflora
crow-foot violet, prairie or crowfoot or larkspur violet, prairie violet, violette pédatifide
arctic yellow violet, European field pansy, northern violet, queen Charlotte twinflower violet, twinflower violet
1–3+, ascending or erect, leafy proximally and distally, glabrous, on caudex from fleshy rhizome.
basal, 2–11, ascending to erect, 5–9-lobed;
stipules linear-lanceolate, margins entire, apex acute;
petiole 3–16 cm, pubescent;
blade similar in width and shape, lobes lanceolate, spatulate, falcate, or linear, 1–7 × 2–8 cm, base truncate to reniform, margins entire, ciliate, apex acute to obtuse, mucronulate, surfaces pubescent, hairs sometimes concentrated on veins.
basal and cauline;
basal: 2–3(–4);
stipules ovate-lanceolate, ovate, or oblong, margins entire, apex ± acute;
petiole 1.5–15 cm, glabrous;
blade broadly reniform to orbiculate, 0.5–4.6 × 0.9–6.4 cm, base cordate, margins crenate to crenate-serrate, ciliate, apex obtuse, rounded, or truncate, rarely with terminal point, abaxial surface sparsely puberulent on veins, adaxial surface glabrate to ± densely puberulent;
cauline similar to basal except: stipules lanceolate, ovate, or oblong, margins entire to erose, apex acute to obtuse;
petiole 0.3–7(–10) cm;
blade sometimes ovate, 0.8–3.7 × 1–4.8 cm.
5–18 cm, glabrous or pubescent.
2–9 cm, usually glabrous.
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm;
petals light to soft reddish violet on both surfaces, lower 3 white basally, dark violet-veined, lateral 2 and lowest usually bearded, lowest 10–25 mm, spur same color as petals, gibbous, 2–3 mm;
style head beardless; cleistogamous flowers on ascending to erect peduncles.
sepals with or without purple stripe on both sides of midvein, narrowly lanceolate to oblong, margins ciliate or eciliate, auricles 0.5–1 mm;
petals deep lemon-yellow on both surfaces, lower 3 and often upper 2 brownish purple-veined, lateral 2 beardless, lowest 6–15 mm, spur yellow to yellowish green, gibbous, 2–2.5 mm;
style head beardless; cleistogamous flowers axillary.
ellipsoid, 10–15 mm, glabrous.
oblong-ovoid, 3–6 mm, glabrous or sparsely pubescent.
beige, mottled to bronze, 1.5–2.5 mm.
purple, sometimes streaked with light and dark brown, 1.5–2.5 mm.
= 54.
Viola pedatifida
Viola biflora
Viola pedatifida was reported historically from, and recently rediscovered in, the Appalachian shale barrens of Virginia (T. Wieboldt, pers. comm.).
Viola pedatifida reportedly hybridizes with V. sororia (= V. ×bernardii Greene).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Varieties 5 (2 in the flora).
Viola biflora may be the most widely distributed species of the genus in the Northern Hemisphere. Although occurring most often in mountainous areas at high elevations, it is known from Alaska near the coast at elevations of ca. 45 m (PNW Herbaria Portal 2010) and has been reported from lowland meadows in Kamchatka (V. B. Baird 1942), and is occasionally found near sea level in exposed rocky habitats on the west coast of the Queen Charlotte Islands (R. A. Calder and R. L. Taylor 1968).
Sometimes described as high latitude, circumpolar, Viola biflora also occurs in mid latitudes north of the equator. It is not truly circumpolar; it does not occur in eastern Canada or in Greenland. The deeply cleft style head sets V. biflora apart from all other species in North America. V. B. Baird (1942) suggested that the occasional presence of two cleistogamous flowers in the axil of the same leaf may account for the name “biflora.”
H. N. Ridley (1930) said that in Europe, deer (Cervus dama Linnaeus and Rangifer tarandus Linnaeus) may play a role in the dispersal of Viola biflora; seeds have been recovered from their droppings.
Nonflowering Viola biflora can be confused with V. renifolia where their ranges overlap in Yukon, Alaska, and Colorado.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Lowest petal 6–10 mm; seeds 1.5–2 mm; sepals usually without purple stripe; Yukon, Alaska, Colorado. | var. biflora |
1. Lowest petal 11–15 mm; seeds 2.5 mm; sepals usually with purple stripe; Queen Charlotte Islands, Vancouver Island, British Columbia. | var. carlottae |
IL Wildflowers
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MI Flora
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