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bird-foot violet

felt-leaf violet, woolly or felt-leaf or feltleaf violet, woolly violet

Habit Plants perennial, acaulescent, not stoloniferous, 5–30 cm; rhizome thick, fleshy. Plants perennial, caulescent, not stoloniferous, 7–10 cm.
Stems

1–3(–5), prostrate or decumbent to erect, leafy proximally and distally, densely white-tomentose, from usually vertical, subligneous rhizome.

Leaves

basal, 4–10, ascending to erect, deeply divided;

stipules linear-lanceolate, margins entire, lacerate, or shallowly divided, apex acute;

petiole 2–12 cm, glabrous or sparsely puberulent;

blade 3–9(–10)-lobed, lobes similar in width and shape, spatulate, lanceolate ± linear, deltate, or ovate, 1–4 × 1–4 cm, base attenuate or broadly cordate to cuneate, margins entire, ciliate or eciliate, apex rounded to usually acute, surfaces usually glabrous, sometimes pubescent on abaxial veins.

basal and cauline;

basal: 1–6;

stipules linear to broadly ovate-oblong, margins entire, sometimes with scattered glandular hairs, apex acute to obtuse;

petiole 2–6 cm, densely white-tomentose;

blade ± oblanceolate to elliptic, 1.5–5 × 1.4–2.1 cm, base attenuate, usually oblique, margins usually entire, rarely crenate distally, ciliate, apex acute to usually obtuse, mucronulate, surfaces densely white-tomentose abaxially, strigose adaxially;

cauline similar to basal except: stipules ovate, lanceolate, oblanceolate, or oblong, margins entire or toothed, densely ciliate with white hairs;

petiole 1.5–3.5 cm;

blade 1.8–4 × 0.6–1.1 cm.

Peduncles

5–12 cm, glabrous or pubescent.

1–4 cm, densely white-tomentose.

Flowers

sepals lanceolate, margins mostly ciliate, at least proximally, auricles 1–2 mm;

petals uniformly light to dark blue-violet on both surfaces or upper 2 darker adaxially, sometimes white, upper and lateral 2 often darker basally, lowest, seldom others, dark violet-veined, all beardless, lowest white basally, 12–24 mm, spur white, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers absent.

sepals lanceolate, margins ciliate-tomentose, auricles 0.5–1 mm;

petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 bearded, lowest 6–11 mm, spur yellow, gibbous, 0.5–1.5 mm;

style head bearded; cleistogamous flowers absent.

Capsules

ellipsoid, 6–10 mm, glabrous.

± spherical, 4–5 mm, densely white-tomentose.

Seeds

beige, mottled to brown, 1.4–3 mm.

brown with lighter brown mottling, 2.5–2.8 mm.

2n

= 56.

= 12.

Viola pedata

Viola tomentosa

Phenology Flowering May–Aug.
Habitat Dry, gravelly places, open ponderosa, Jeffrey, lodgepole pine forests
Elevation 1500–2000 m (4900–6600 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; ON
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from FNA
CA
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[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola tomentosa occurs in El Dorado, Nevada, Placer, Plumas, and Sierra counties. M. S. Baker (1949) reported that nearly every leaf axil of V. tomentosa produces a flower bud and that these buds produce chasmogamous flowers late in season instead of cleistogamous flowers, as do other members of the V. nuttallii complex.

Viola tomentosa hybridizes with V. purpurea; the hybrids appeared sterile (M. S. Baker 1949). J. Clausen (1964) reported a putative hybrid between V. tomentosa and V. sheltonii from one location in Sierra County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades 7–9-lobed, lobes spatulate, lanceolate, or ± linear, sometimes with narrowly deltate to falcate appendages toward apex.
var. pedata
1. Leaf blades 3–5(–10)-lobed, lobes deltate or ovate.
var. ranunculifolia
Source FNA vol. 6, p. 143. FNA vol. 6, p. 160.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. pedata var. pedata, V. pedata var. ranunculifolia
Name authority Linnaeus: Sp. Pl. 2: 933. (1753) M. S. Baker & J. C. Clausen: Leafl. W. Bot. 5: 142. (1949)
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