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bird-foot violet

Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet

Habit Plants perennial, acaulescent, not stoloniferous, 5–30 cm; rhizome thick, fleshy. Plants perennial, caulescent, not stoloniferous, 3–20 cm.
Stems

1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex.

Leaves

basal, 4–10, ascending to erect, deeply divided;

stipules linear-lanceolate, margins entire, lacerate, or shallowly divided, apex acute;

petiole 2–12 cm, glabrous or sparsely puberulent;

blade 3–9(–10)-lobed, lobes similar in width and shape, spatulate, lanceolate ± linear, deltate, or ovate, 1–4 × 1–4 cm, base attenuate or broadly cordate to cuneate, margins entire, ciliate or eciliate, apex rounded to usually acute, surfaces usually glabrous, sometimes pubescent on abaxial veins.

basal and cauline;

basal: 1–6, bipinnately compound, leaflets 3–5;

stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate;

petiole 5–6.8 cm, glabrous or puberulent;

blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;

cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute;

petiole 0.9–4 cm;

blade 1.1–4.1 × 1–3.6 cm.

Peduncles

5–12 cm, glabrous or pubescent.

2–12.5 cm, glabrous or puberulent.

Flowers

sepals lanceolate, margins mostly ciliate, at least proximally, auricles 1–2 mm;

petals uniformly light to dark blue-violet on both surfaces or upper 2 darker adaxially, sometimes white, upper and lateral 2 often darker basally, lowest, seldom others, dark violet-veined, all beardless, lowest white basally, 12–24 mm, spur white, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers absent.

sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;

petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm;

style head bearded; cleistogamous flowers absent.

Capsules

ellipsoid, 6–10 mm, glabrous.

spherical to oblong, 5–12 mm, glabrous.

Seeds

beige, mottled to brown, 1.4–3 mm.

light brown, 2.8–3.3 mm.

2n

= 56.

= 24, 48.

Viola pedata

Viola douglasii

Phenology Flowering Feb–Jul.
Habitat Vernally moist grassy slopes and flats, often serpentine soil (except Oregon)
Elevation 20–2300 m (100–7500 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; ON
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from FNA
CA; OR; Mexico (Baja California)
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Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades 7–9-lobed, lobes spatulate, lanceolate, or ± linear, sometimes with narrowly deltate to falcate appendages toward apex.
var. pedata
1. Leaf blades 3–5(–10)-lobed, lobes deltate or ovate.
var. ranunculifolia
Source FNA vol. 6, p. 143. FNA vol. 6, p. 129.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. pedata var. pedata, V. pedata var. ranunculifolia
Synonyms V. chrysantha
Name authority Linnaeus: Sp. Pl. 2: 933. (1753) Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841)
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