Viola palmata
Viola tomentosa
early blue violet, three-lobe violet, three-lobed violet, trilobed violet, wood violet
felt-leaf violet, woolly or felt-leaf or feltleaf violet, woolly violet
1–3(–5), prostrate or decumbent to erect, leafy proximally and distally, densely white-tomentose, from usually vertical, subligneous rhizome.
basal, 2–3, ascending to erect;
stipules linear-lanceolate, margins entire, apex acute;
petiole 1–20 cm, glabrous or pubescent;
earliest leaf blades unlobed, mid-season blades 3–9-lobed, lobes sometimes appearing petiolate and sometimes further lobed;
earliest leaf blades reniform to ovate, mid-season blades with middle lobes usually ovate or elliptic to widely obovate, sometimes narrowly elliptic, narrowly ovate, lanceolate, or spatulate to narrowly obovate, lateral lobes elliptic, obdeltate, or spatulate to falcate, smaller lobes similar, 1–14 × 1–10 cm, base truncate to cordate, margins serrate or entire, usually ciliate, apex acute, rounded, blunt, or obtuse, surfaces glabrous or pubescent throughout or along veins.
basal and cauline;
basal: 1–6;
stipules linear to broadly ovate-oblong, margins entire, sometimes with scattered glandular hairs, apex acute to obtuse;
petiole 2–6 cm, densely white-tomentose;
blade ± oblanceolate to elliptic, 1.5–5 × 1.4–2.1 cm, base attenuate, usually oblique, margins usually entire, rarely crenate distally, ciliate, apex acute to usually obtuse, mucronulate, surfaces densely white-tomentose abaxially, strigose adaxially;
cauline similar to basal except: stipules ovate, lanceolate, oblanceolate, or oblong, margins entire or toothed, densely ciliate with white hairs;
petiole 1.5–3.5 cm;
blade 1.8–4 × 0.6–1.1 cm.
3–13 cm, glabrous or pubescent.
1–4 cm, densely white-tomentose.
sepals lanceolate, margins ciliate or eciliate, auricles 0.5–1 mm;
petals violet on both surfaces, lower 3 white basally, lower 3 and upper 2 sometimes purple-veined, lateral 2 bearded, spur sometimes bearded, lowest 15–25 mm, spur white, gibbous, 2–3 mm;
style head beardless; cleistogamous flowers on prostrate to ascending peduncles.
sepals lanceolate, margins ciliate-tomentose, auricles 0.5–1 mm;
petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 bearded, lowest 6–11 mm, spur yellow, gibbous, 0.5–1.5 mm;
style head bearded; cleistogamous flowers absent.
ellipsoid, 5–15 mm, glabrous.
± spherical, 4–5 mm, densely white-tomentose.
beige, mottled to bronze, 1.5–2.5 mm.
brown with lighter brown mottling, 2.5–2.8 mm.
= 54.
= 12.
Viola palmata
Viola tomentosa
N. L. Gil-Ad (1997) presented evidence based on seed coat micromorphology that some plants known as Viola palmata are hybrids between plants with lobed and unlobed leaves and believed the type specimen to be hybrid in origin. He chose not to recognize the name V. palmata, choosing instead to recognize V. triloba. If the type specimen does represent a hybrid, Gil-Ad would be correct in recognizing V. triloba. Because the purity of the type specimen cannot be ascertained, we consider V. palmata the most appropriate name for this taxon. We acknowledge the presence of hybrids between lobed and unlobed plants; such hybrids obscure lines of demarcation. Some herbarium specimens may represent such hybrids; we feel most do not.
L. E. McKinney (1992) described the nomenclatural history of Viola palmata. Homophylly versus heterophylly differentiates V. palmata from V. subsinuata, the homophyllous violet most often called V. palmata. We know that the taxon described by Linnaeus was heterophyllous, with some undivided leaf blades.
Varieties 2 (2 in the flora).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Viola tomentosa occurs in El Dorado, Nevada, Placer, Plumas, and Sierra counties. M. S. Baker (1949) reported that nearly every leaf axil of V. tomentosa produces a flower bud and that these buds produce chasmogamous flowers late in season instead of cleistogamous flowers, as do other members of the V. nuttallii complex.
Viola tomentosa hybridizes with V. purpurea; the hybrids appeared sterile (M. S. Baker 1949). J. Clausen (1964) reported a putative hybrid between V. tomentosa and V. sheltonii from one location in Sierra County.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Petioles, leaf surfaces, and peduncles usually pubescent, rarely glabrous; apex of middle leaf lobe acute; plants of dry to mesic habitats. | var. palmata |
1. Petioles, leaf surfaces, and peduncles usually glabrous, rarely pubescent; apex of middle leaf lobe rounded, blunt, or obtuse; plants of wet ground. | var. heterophylla |
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