Viola odorata |
Viola utahensis |
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English violet, garden violet, sweet blue violet, sweet violet, violette odorante |
Utah violet |
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Habit | Plants perennial, acaulescent, stoloniferous, 4–12 cm; stolons green, often rooting at nodes and forming leafy rosettes; rooted rosettes often develop into erect, rhizomatous caudex from which new stolons are produced; rhizome thick, fleshy. | Plants perennial, caulescent, not stoloniferous, 5–20 cm. |
Stems | 1–4, prostrate, decumbent, or erect, leafy proximally and distally, glabrate or sparsely puberulent, on caudex from subligneous rhizome. |
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Leaves | basal (and from stolons), 5–10, ascending to erect; stipules lanceolate to linear-lanceolate, margins fimbriate, projections gland-tipped, apex acute; petiole 2–17 cm, puberulent; blade unlobed, ovate to orbiculate, 1.5–7 × 1.5–5 cm, base cordate, margins crenate, ciliate, apex obtuse to rounded, surfaces puberulent. |
basal and cauline; basal: 1–3; stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire or laciniate with glandular hairs, apex of each wing free, acute, sometimes divided into several filiform processes; petiole 2–8(–12) cm, glabrate or sparsely puberulent; blade usually broadly ovate, sometimes ± orbiculate, 1–4.5 × 0.7–3.4 cm, base usually attenuate, rarely truncate or subcordate, margins ± coarsely crenate-serrate, ciliate, apex obtuse, surfaces glabrate or sparsely puberulent; cauline similar to basal except: stipules lanceolate or linear-lanceolate to linear, margins ± entire or lacerate to laciniate with sparse gland-tipped projections, apex acute to acuminate, often divided into 2–4 filiform processes; petiole 3–8.7 cm; blade ovate to oblong-ovate or oblong-lanceolate, 0.7–1.6 × 2.6–3.7 cm, margins shallowly serrate, irregularly crenate, or ± entire, apex acute to obtuse. |
Peduncles | 4–15 cm, puberulent. |
4–10 cm, glabrate or sparsely puberulent. |
Flowers | sepals narrow to broadly lanceolate, margins ciliate, auricles 3–4 mm; petals deep to pale blue-violet, pale blue, or white on both surfaces, usually white basally, lateral 2 sparsely to densely bearded, lowest usually purple-veined, 12–22 mm, spur usually same color as petals, elongated, 5–7 mm; style head beardless; cleistogamous flowers on prostrate or ascending peduncles. |
sepals lanceolate, margins ciliate or eciliate, auricles 1–1.5 mm; petals deep lemon-yellow adaxially, upper 2 and sometimes lateral 2 reddish brown to brownish purple abaxially, lower 3 and sometimes upper 2 dark brown-veined, lateral 2 sparsely bearded, lowest 10–13 mm, spur yellow, gibbous, 0.5–1.5 mm; style head bearded; cleistogamous flowers axillary. |
Capsules | sometimes purple-flecked, ovoid, 5–8 mm, puberulent. |
± spherical, 6–7 mm, glabrous or finely puberulent. |
Seeds | brown, 3–4 mm. |
brown, ca. 3 mm. |
2n | = 20. |
= 24. |
Viola odorata |
Viola utahensis |
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Phenology | Flowering Jan–May. | Flowering May. |
Habitat | Lawns, roadsides, clearings, riparian habitats, parks, urban areas | Open sagebrush areas, semidesert slopes, grasslands, open forests |
Elevation | 0–1700 m (0–5600 ft) | 1200–2600 m (3900–8500 ft) |
Distribution |
CA; CT; ID; IL; MA; ME; MI; NC; NY; OH; OR; PA; RI; UT; WA; WI; BC; NS; ON; QC; Eurasia [Introduced in North America; introduced also in Australia]
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CO; ID; NV; UT; WY
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Discussion | Viola odorata occurs in small colonies; individual plants are interconnected by stolons. The flowers of Viola odorata are noted for their fragrance; some plants are more fragrant than others. It is native to Eurasia and assumed to be introduced in North America where it is usually found in areas associated with human habitation, including parks, lawns, and roadsides. A substantial industry revolved around the commercial production of violets in England, France, and the United States from prior to 1895 and into the 1900s (R. E. Coombs 2003). Viola odorata is sometimes found in remote locations not easily explained by anthropogenic influence, for example, Clearwater Mountains, Idaho. It is available through the nursery trade and is cultivated as a garden plant and occasionally reported as an escape. Viola odorata is grown in southern France for essential oils used in perfumes, flavorings, and toiletries, and also for the production of the sweet, violet-colored liqueur called parfait amour (V. H. Heywood 1978). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola utahensis occurs only in areas where its closest relatives, V. vallicola and V. purpurea var. venosa, are also found and it appears to have originated from the crossing of those taxa followed by a doubling of the chromosomes (M. S. Baker 1949). The elaiosome of V. utahensis seeds has the same general appearance as those of other members of V. purpurea complex (except V. purpurea var. venosa), covering about one-third the length of the seed. G. Davidse (1976) found that tetraploid members of Clausen’s “subsect.” Purpurea (V. utahensis) have serrate leaf margins; diploid plants (V. purpurea) have crenate leaf margins. He also found that the size of V. utahensis plants is correlated with elevation, the largest plants at low elevations and the smallest at higher elevations. G. Davidse (1976) reported putative hybrids between Viola utahensis and V. praemorsa subsp. major (= var. linguifolia), V. purpurea var. venosa, and V. beckwithii. Observed pollinators of Viola utahensis in Utah include the bees Apis mellifera Linnaeus and Anthophora ursina Cresson (G. Davidse 1976). In other populations, flies in the genera Eristalis Latreille and Bombylius Linnaeus were observed indiscriminately visiting the yellow-flowered species V. praemorsa subsp. major, V. utahensis, and V. vallicola (Davidse). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 140. | FNA vol. 6, p. 162. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Name authority | Linnaeus: Sp. Pl. 2: 934. (1753) | M. S. Baker & J. C. Clausen: Leafl. W. Bot. 5: 145. (1949) |
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