Viola nephrophylla |
Viola purpurea |
|||||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
kidney-leaf violet, Leconte violet, northern bog violet, violette néphrophylle |
goosefoot violet, mountain violet, pine violet, purple-mark yellow violet |
|||||||||||||||||||||||||
Habit | Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age. | Plants perennial, caulescent, not stoloniferous, 1.5–25 cm. | ||||||||||||||||||||||||
Stems | 1–5(–7), decumbent or spreading to erect, leafy proximally and distally, ± glabrous, puberulent, canescent, or tomentose, on caudex from subligneous rhizome. |
|||||||||||||||||||||||||
Leaves | basal, 4–7, ascending to erect; stipules lanceolate, margins entire or fimbriate, apex acute; petiole 2–25 cm, usually glabrous; blade usually grayish green or purplish green abaxially, unlobed, ovate, reniform, or broadly reniform to orbiculate, 1–7 × 1–7 cm, somewhat fleshy, base broadly cordate or reniform to ± truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually glabrous, rarely sparsely pubescent. |
basal and cauline; basal: 1–6; stipules adnate to petiole, forming 2 linear, membranous wings, wing margins entire or laciniate, each wing with lanceolate to ± deltate projection, margins entire or laciniate, apex acute to long-acuminate; petiole 1.8–14.5 cm, puberulent to tomentose; blade purplish, purple-tinted, or gray-green abaxially, gray, green, or gray-green adaxially, sometimes shiny adaxially, ovate, orbiculate, oblong, deltate, or lanceolate, 0.8–5.3 × 0.4–4.1 cm, often fleshy, base cordate, subcordate, truncate, or attenuate, oblique or not, margins usually ± crenate, serrate, dentate, or coarsely or irregularly repand-dentate, sometimes entire, usually ciliate, apex acute to obtuse or rounded, surfaces glabrous, puberulent, or tomentose; cauline similar to basal except: stipules linear, lanceolate, oblanceolate, or ± oblong to ovate, margins entire, lacerate, or laciniate, usually ciliate, apex sometimes divided into 2–3 filiform processes or obtuse; petiole 0.3–19.7 cm, glabrous or puberulent; blade ovate, oblong, elliptic, deltate, or lanceolate, 0.9–5.2 × 0.2–2.9 cm, length 0.8–7.1 times width, margins crenate, serrate, dentate, repand-denticulate, undulate-denticulate, sinuate, undulate, or entire, abaxial surface puberulent, canescent, or tomentose, adaxial surface glabrous, sparsely pubescent, puberulent, canescent, or tomentose. |
||||||||||||||||||||||||
Peduncles | 3–25 cm, usually glabrous. |
|||||||||||||||||||||||||
Flowers | sepals ovate, margins usually eciliate, auricles 1–2 mm; petals deep bluish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, upper 2 sometimes sparsely bearded, lowest densely bearded or beardless, 10–28 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on erect to ascending peduncles. |
|||||||||||||||||||||||||
Capsules | ovoid, 5–10 mm, glabrous. |
ovoid to ± spherical, 4–7 mm, puberulent. |
||||||||||||||||||||||||
Seeds | beige to brown or dark brown, 1.5–2.5 mm. |
light to dark brown or mottled gray and brown, 2–3.1 mm. |
||||||||||||||||||||||||
2n | = 54. |
= 12. |
||||||||||||||||||||||||
Viola nephrophylla |
Viola purpurea |
|||||||||||||||||||||||||
Phenology | Flowering Jan–Sep. | |||||||||||||||||||||||||
Habitat | Wet habitats in saturated soil in prairies, pastures, bogs, fens, sedge meadows, rocky shores of lakes and streams, limestone outcrops, gravelly calcareous stream beds | |||||||||||||||||||||||||
Elevation | 100–3000 m (300–9800 ft) | |||||||||||||||||||||||||
Distribution |
AR; AZ; CA; CO; CT; IA; ID; IL; KS; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OH; OR; PA; SD; TX; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT
|
AZ; CA; CO; ID; MT; NV; OR; UT; WA; WY; BC; nw Mexico
|
||||||||||||||||||||||||
Discussion | Viola nephrophylla has the widest distribution in North America of any of the acaulescent blue violets. There is some question whether the species we now know as Viola nephrophylla fits with the protologue and type specimen(s). McKinney examined the type designated by Greene; it appears that what most botanists have called V. nephrophylla may be an undescribed taxon. N. L. Gil-Ad (1997) believed Greene’s type contained plants that were genetically impure based on seed coat micromorphology and suggested the specimens might be a mixture of both orthospecies and introgressants. Viola mccabeiana was described by M. S. Baker in 1940 from collections made in British Columbia. Later, Baker (1949b) discussed the reasons why the name V. mccabeiana needed to be abandoned. C. L. Hitchcock et al. (1955–1969, vol. 3) treated V. mccabeiana as synonymous with V. nephrophylla var. cognata. While Viola mccabeiana is recognized in Canada (L. Brouillet et al., http://data.canadensys.net/vascan/), we believe it is best included within V. nephrophylla. Viola nephrophylla reportedly hybridizes with V. cucullata (= V. ×insessa House), V. sororia var. sororia (= V. ×napae House), and V. affinis (= V. ×subaffinis House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 7 (7 in the flora). Varieties of Viola purpurea are variable and intergrade. All are found in California; six occur in other western states, one in Mexico, and one in British Columbia. Mature plants are needed for determination. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||||||||||||||
Key |
|
|||||||||||||||||||||||||
Source | FNA vol. 6, p. 139. | FNA vol. 6, p. 150. | ||||||||||||||||||||||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||
Synonyms | V. arizonica, V. austiniae, V. cognata, V. crassula, V. galacifolia, V. lunellii, V. mccabeiana, V. nephrophylla var. arizonica, V. nephrophylla var. cognata, V. peramoena, V. pratincola, V. prionosepala, V. retusa, V. subjuncta, V. vagula | |||||||||||||||||||||||||
Name authority | Greene: Pittonia 3: 144. (1896) | Kellogg: Pacific (San Francisco), 2 Feb. 1855: unnumb. (1855) | ||||||||||||||||||||||||
Web links |
|
|