Viola nephrophylla |
Viola pedatifida |
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kidney-leaf violet, Leconte violet, northern bog violet, violette néphrophylle |
crow-foot violet, prairie or crowfoot or larkspur violet, prairie violet, violette pédatifide |
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Habit | Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age. | Plants perennial, acaulescent, not stoloniferous, 5–30 cm; rhizome thick, fleshy. |
Leaves | basal, 4–7, ascending to erect; stipules lanceolate, margins entire or fimbriate, apex acute; petiole 2–25 cm, usually glabrous; blade usually grayish green or purplish green abaxially, unlobed, ovate, reniform, or broadly reniform to orbiculate, 1–7 × 1–7 cm, somewhat fleshy, base broadly cordate or reniform to ± truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually glabrous, rarely sparsely pubescent. |
basal, 2–11, ascending to erect, 5–9-lobed; stipules linear-lanceolate, margins entire, apex acute; petiole 3–16 cm, pubescent; blade similar in width and shape, lobes lanceolate, spatulate, falcate, or linear, 1–7 × 2–8 cm, base truncate to reniform, margins entire, ciliate, apex acute to obtuse, mucronulate, surfaces pubescent, hairs sometimes concentrated on veins. |
Peduncles | 3–25 cm, usually glabrous. |
5–18 cm, glabrous or pubescent. |
Flowers | sepals ovate, margins usually eciliate, auricles 1–2 mm; petals deep bluish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, upper 2 sometimes sparsely bearded, lowest densely bearded or beardless, 10–28 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on erect to ascending peduncles. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to soft reddish violet on both surfaces, lower 3 white basally, dark violet-veined, lateral 2 and lowest usually bearded, lowest 10–25 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on ascending to erect peduncles. |
Capsules | ovoid, 5–10 mm, glabrous. |
ellipsoid, 10–15 mm, glabrous. |
Seeds | beige to brown or dark brown, 1.5–2.5 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
2n | = 54. |
= 54. |
Viola nephrophylla |
Viola pedatifida |
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Phenology | Flowering Jan–Sep. | Flowering Mar–Jun. |
Habitat | Wet habitats in saturated soil in prairies, pastures, bogs, fens, sedge meadows, rocky shores of lakes and streams, limestone outcrops, gravelly calcareous stream beds | Prairies, grasslands, disturbed ground, dry gravelly hills |
Elevation | 100–3000 m (300–9800 ft) | 500–1000 m (1600–3300 ft) |
Distribution |
AR; AZ; CA; CO; CT; IA; ID; IL; KS; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OH; OR; PA; SD; TX; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT
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AR; AZ; CO; IA; IL; IN; KS; MI; MN; MO; MT; ND; NE; NM; OH; OK; SD; VA; WI; WY; AB; MB; ON; SK
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Discussion | Viola nephrophylla has the widest distribution in North America of any of the acaulescent blue violets. There is some question whether the species we now know as Viola nephrophylla fits with the protologue and type specimen(s). McKinney examined the type designated by Greene; it appears that what most botanists have called V. nephrophylla may be an undescribed taxon. N. L. Gil-Ad (1997) believed Greene’s type contained plants that were genetically impure based on seed coat micromorphology and suggested the specimens might be a mixture of both orthospecies and introgressants. Viola mccabeiana was described by M. S. Baker in 1940 from collections made in British Columbia. Later, Baker (1949b) discussed the reasons why the name V. mccabeiana needed to be abandoned. C. L. Hitchcock et al. (1955–1969, vol. 3) treated V. mccabeiana as synonymous with V. nephrophylla var. cognata. While Viola mccabeiana is recognized in Canada (L. Brouillet et al., http://data.canadensys.net/vascan/), we believe it is best included within V. nephrophylla. Viola nephrophylla reportedly hybridizes with V. cucullata (= V. ×insessa House), V. sororia var. sororia (= V. ×napae House), and V. affinis (= V. ×subaffinis House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola pedatifida was reported historically from, and recently rediscovered in, the Appalachian shale barrens of Virginia (T. Wieboldt, pers. comm.). Viola pedatifida reportedly hybridizes with V. sororia (= V. ×bernardii Greene). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 139. | FNA vol. 6, p. 144. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. arizonica, V. austiniae, V. cognata, V. crassula, V. galacifolia, V. lunellii, V. mccabeiana, V. nephrophylla var. arizonica, V. nephrophylla var. cognata, V. peramoena, V. pratincola, V. prionosepala, V. retusa, V. subjuncta, V. vagula | V. delphiniifolia, V. palmata var. pedatifida |
Name authority | Greene: Pittonia 3: 144. (1896) | G. Don: Gen. Hist. 1: 320. (1831) |
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