Viola nephrophylla |
Viola palmata |
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kidney-leaf violet, Leconte violet, northern bog violet, violette néphrophylle |
early blue violet, three-lobe violet, three-lobed violet, trilobed violet, wood violet |
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Habit | Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age. | Plants perennial, acaulescent, not stoloniferous, 6–50 cm; rhizome thick, fleshy. | ||||
Leaves | basal, 4–7, ascending to erect; stipules lanceolate, margins entire or fimbriate, apex acute; petiole 2–25 cm, usually glabrous; blade usually grayish green or purplish green abaxially, unlobed, ovate, reniform, or broadly reniform to orbiculate, 1–7 × 1–7 cm, somewhat fleshy, base broadly cordate or reniform to ± truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually glabrous, rarely sparsely pubescent. |
basal, 2–3, ascending to erect; stipules linear-lanceolate, margins entire, apex acute; petiole 1–20 cm, glabrous or pubescent; earliest leaf blades unlobed, mid-season blades 3–9-lobed, lobes sometimes appearing petiolate and sometimes further lobed; earliest leaf blades reniform to ovate, mid-season blades with middle lobes usually ovate or elliptic to widely obovate, sometimes narrowly elliptic, narrowly ovate, lanceolate, or spatulate to narrowly obovate, lateral lobes elliptic, obdeltate, or spatulate to falcate, smaller lobes similar, 1–14 × 1–10 cm, base truncate to cordate, margins serrate or entire, usually ciliate, apex acute, rounded, blunt, or obtuse, surfaces glabrous or pubescent throughout or along veins. |
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Peduncles | 3–25 cm, usually glabrous. |
3–13 cm, glabrous or pubescent. |
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Flowers | sepals ovate, margins usually eciliate, auricles 1–2 mm; petals deep bluish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, upper 2 sometimes sparsely bearded, lowest densely bearded or beardless, 10–28 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on erect to ascending peduncles. |
sepals lanceolate, margins ciliate or eciliate, auricles 0.5–1 mm; petals violet on both surfaces, lower 3 white basally, lower 3 and upper 2 sometimes purple-veined, lateral 2 bearded, spur sometimes bearded, lowest 15–25 mm, spur white, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
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Capsules | ovoid, 5–10 mm, glabrous. |
ellipsoid, 5–15 mm, glabrous. |
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Seeds | beige to brown or dark brown, 1.5–2.5 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
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2n | = 54. |
= 54. |
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Viola nephrophylla |
Viola palmata |
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Phenology | Flowering Jan–Sep. | |||||
Habitat | Wet habitats in saturated soil in prairies, pastures, bogs, fens, sedge meadows, rocky shores of lakes and streams, limestone outcrops, gravelly calcareous stream beds | |||||
Elevation | 100–3000 m (300–9800 ft) | |||||
Distribution |
AR; AZ; CA; CO; CT; IA; ID; IL; KS; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OH; OR; PA; SD; TX; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT
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AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; ND; NE; NJ; NM; OH; OK; PA; RI; SC; TN; TX; VA; VT; WV; ON
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Discussion | Viola nephrophylla has the widest distribution in North America of any of the acaulescent blue violets. There is some question whether the species we now know as Viola nephrophylla fits with the protologue and type specimen(s). McKinney examined the type designated by Greene; it appears that what most botanists have called V. nephrophylla may be an undescribed taxon. N. L. Gil-Ad (1997) believed Greene’s type contained plants that were genetically impure based on seed coat micromorphology and suggested the specimens might be a mixture of both orthospecies and introgressants. Viola mccabeiana was described by M. S. Baker in 1940 from collections made in British Columbia. Later, Baker (1949b) discussed the reasons why the name V. mccabeiana needed to be abandoned. C. L. Hitchcock et al. (1955–1969, vol. 3) treated V. mccabeiana as synonymous with V. nephrophylla var. cognata. While Viola mccabeiana is recognized in Canada (L. Brouillet et al., http://data.canadensys.net/vascan/), we believe it is best included within V. nephrophylla. Viola nephrophylla reportedly hybridizes with V. cucullata (= V. ×insessa House), V. sororia var. sororia (= V. ×napae House), and V. affinis (= V. ×subaffinis House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
N. L. Gil-Ad (1997) presented evidence based on seed coat micromorphology that some plants known as Viola palmata are hybrids between plants with lobed and unlobed leaves and believed the type specimen to be hybrid in origin. He chose not to recognize the name V. palmata, choosing instead to recognize V. triloba. If the type specimen does represent a hybrid, Gil-Ad would be correct in recognizing V. triloba. Because the purity of the type specimen cannot be ascertained, we consider V. palmata the most appropriate name for this taxon. We acknowledge the presence of hybrids between lobed and unlobed plants; such hybrids obscure lines of demarcation. Some herbarium specimens may represent such hybrids; we feel most do not. L. E. McKinney (1992) described the nomenclatural history of Viola palmata. Homophylly versus heterophylly differentiates V. palmata from V. subsinuata, the homophyllous violet most often called V. palmata. We know that the taxon described by Linnaeus was heterophyllous, with some undivided leaf blades. Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 139. | FNA vol. 6, p. 141. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. arizonica, V. austiniae, V. cognata, V. crassula, V. galacifolia, V. lunellii, V. mccabeiana, V. nephrophylla var. arizonica, V. nephrophylla var. cognata, V. peramoena, V. pratincola, V. prionosepala, V. retusa, V. subjuncta, V. vagula | V. cucullata var. palmata | ||||
Name authority | Greene: Pittonia 3: 144. (1896) | Linnaeus: Sp. Pl. 2: 933. (1753) | ||||
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