Viola nephrophylla
Viola orbiculata
kidney-leaf violet, Leconte violet, northern bog violet, violette néphrophylle
darkwoods violet, evergreen yellow violet, round-leaf or round-leaf wood or western round-leaf or evergreen violet, round-leaf violet, violette orbiculaire
1–5, prostrate or erect, leafy proximally and distally, usually glabrous, from current and/or previous year’s growth, on usually vertical, fleshy rhizome.
basal, 4–7, ascending to erect;
stipules lanceolate, margins entire or fimbriate, apex acute;
petiole 2–25 cm, usually glabrous;
blade usually grayish green or purplish green abaxially, unlobed, ovate, reniform, or broadly reniform to orbiculate, 1–7 × 1–7 cm, somewhat fleshy, base broadly cordate or reniform to ± truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually glabrous, rarely sparsely pubescent.
basal and cauline;
basal: 3–11;
stipules deltate to lanceolate, margins entire, apex acute to acuminate;
petiole 2.1–10.5 cm, glabrous or pubescent;
blade usually orbiculate to broadly ovate, sometimes reniform, 1.4–5.3 × 1.4–5.3 cm, base cordate, margins crenate to serrulate-crenulate, eciliate or sparingly ciliate, apex usually obtuse, rarely acute, abaxial surface usually glabrous, adaxial surface sparsely pubescent;
cauline similar to basal except: stipules lanceolate to ovate, margins usually entire, ciliate;
petiole 0.7–1 cm, glabrous or pubescent;
blade 1.2–2 × 1.1–1.4 cm.
3–25 cm, usually glabrous.
2.7–5.6 cm, glabrous.
sepals ovate, margins usually eciliate, auricles 1–2 mm;
petals deep bluish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, upper 2 sometimes sparsely bearded, lowest densely bearded or beardless, 10–28 mm, spur same color as petals, gibbous, 2–3 mm;
style head beardless; cleistogamous flowers on erect to ascending peduncles.
sepals narrow to broadly lanceolate, margins eciliate, auricles 0.5–1.5 mm;
upper 2 and lateral 2 petals deep lemon-yellow on both surfaces, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 usually bearded, lowest 8–17 mm, spur usually yellow, sometimes whitish, gibbous, 1.5–2.5 mm;
style head bearded; cleistogamous flowers axillary.
ovoid, 5–10 mm, glabrous.
ellipsoid, 6–8 mm, glabrous.
beige to brown or dark brown, 1.5–2.5 mm.
light to dark brown, sometimes mottled with white, 2–2.3 mm.
= 54.
= 24.
Viola nephrophylla
Viola orbiculata
Viola nephrophylla has the widest distribution in North America of any of the acaulescent blue violets.
There is some question whether the species we now know as Viola nephrophylla fits with the protologue and type specimen(s). McKinney examined the type designated by Greene; it appears that what most botanists have called V. nephrophylla may be an undescribed taxon. N. L. Gil-Ad (1997) believed Greene’s type contained plants that were genetically impure based on seed coat micromorphology and suggested the specimens might be a mixture of both orthospecies and introgressants.
Viola mccabeiana was described by M. S. Baker in 1940 from collections made in British Columbia. Later, Baker (1949b) discussed the reasons why the name V. mccabeiana needed to be abandoned. C. L. Hitchcock et al. (1955–1969, vol. 3) treated V. mccabeiana as synonymous with V. nephrophylla var. cognata. While Viola mccabeiana is recognized in Canada (L. Brouillet et al., http://data.canadensys.net/vascan/), we believe it is best included within V. nephrophylla.
Viola nephrophylla reportedly hybridizes with V. cucullata (= V. ×insessa House), V. sororia var. sororia (= V. ×napae House), and V. affinis (= V. ×subaffinis House).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
During winter, the basal leaves of Viola orbiculata are appressed to the ground under the weight of snow. The leaves overwinter and are often partly or entirely green after the snow has melted. Mature plants possess 3–11 basal leaves, some derived from previous years and others from the current year’s growth. Occasional plants are found with short, thin stems rooted at the tip and producing a rosette of leaves and stems, showing the close relationship with V. sempervirens.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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