Viola nephrophylla |
Viola ocellata |
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kidney-leaf violet, Leconte violet, northern bog violet, violette néphrophylle |
pinto violet, two-eyed violet, western heart's ease, western heart's ease or two-eyed or southern two-eyed violet |
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Habit | Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age. | Plants perennial, caulescent, not stoloniferous, 2–37 cm. |
Stems | 1–3, ascending to erect, ± glabrous or usually puberulent, on caudex from shallow, fleshy rhizome or deep-seated caudex with fleshy roots. |
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Leaves | basal, 4–7, ascending to erect; stipules lanceolate, margins entire or fimbriate, apex acute; petiole 2–25 cm, usually glabrous; blade usually grayish green or purplish green abaxially, unlobed, ovate, reniform, or broadly reniform to orbiculate, 1–7 × 1–7 cm, somewhat fleshy, base broadly cordate or reniform to ± truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually glabrous, rarely sparsely pubescent. |
basal and cauline; basal: 1–6; stipules deltate, margins laciniate, apex usually long-acuminate; petiole 3.7–10 cm, puberulent; blade ovate, deltate, or subreniform, 1–6 × 1.2–4 cm, base usually cordate to subcordate, margins crenate, usually ciliate, apex acute, mucronulate, surfaces usually sparsely puberulent, glabrous early; cauline similar to basal except: stipules lanceolate, margins ±fimbriate, sometimes stipitate-glandular, sometimes entire, erose, or ± laciniate; petiole 0.4–9 cm; blade ovate to deltate, 1.6–4.4 × 1.1–3.6 cm, base subcordate to truncate, margins crenate to ± serrate, ciliate, apex acute to obtuse. |
Peduncles | 3–25 cm, usually glabrous. |
1–10 cm, puberulent. |
Flowers | sepals ovate, margins usually eciliate, auricles 1–2 mm; petals deep bluish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, upper 2 sometimes sparsely bearded, lowest densely bearded or beardless, 10–28 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on erect to ascending peduncles. |
sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals white adaxially, upper 2 and sometimes lower 3 deep reddish violet abaxially, all with yellow area basally, lateral 2 with purple patch basally distal to smaller yellow area, bearded, lowest with yellow patch basally, purple-veined, 5–15 mm, spur yellow or greenish, gibbous, 1–2.5 mm; style head bearded; cleistogamous flowers axillary. |
Capsules | ovoid, 5–10 mm, glabrous. |
spherical to spherical-ovoid, 5–11 mm, minutely scabrous. |
Seeds | beige to brown or dark brown, 1.5–2.5 mm. |
brown-purple, ca. 2 mm. |
2n | = 54. |
= 12. |
Viola nephrophylla |
Viola ocellata |
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Phenology | Flowering Jan–Sep. | Flowering Mar–Jul. |
Habitat | Wet habitats in saturated soil in prairies, pastures, bogs, fens, sedge meadows, rocky shores of lakes and streams, limestone outcrops, gravelly calcareous stream beds | Rocky areas, grassy banks, thickets, often on serpentine soil |
Elevation | 100–3000 m (300–9800 ft) | 100–1100 m (300–3600 ft) |
Distribution |
AR; AZ; CA; CO; CT; IA; ID; IL; KS; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OH; OR; PA; SD; TX; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT
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CA; OR
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Discussion | Viola nephrophylla has the widest distribution in North America of any of the acaulescent blue violets. There is some question whether the species we now know as Viola nephrophylla fits with the protologue and type specimen(s). McKinney examined the type designated by Greene; it appears that what most botanists have called V. nephrophylla may be an undescribed taxon. N. L. Gil-Ad (1997) believed Greene’s type contained plants that were genetically impure based on seed coat micromorphology and suggested the specimens might be a mixture of both orthospecies and introgressants. Viola mccabeiana was described by M. S. Baker in 1940 from collections made in British Columbia. Later, Baker (1949b) discussed the reasons why the name V. mccabeiana needed to be abandoned. C. L. Hitchcock et al. (1955–1969, vol. 3) treated V. mccabeiana as synonymous with V. nephrophylla var. cognata. While Viola mccabeiana is recognized in Canada (L. Brouillet et al., http://data.canadensys.net/vascan/), we believe it is best included within V. nephrophylla. Viola nephrophylla reportedly hybridizes with V. cucullata (= V. ×insessa House), V. sororia var. sororia (= V. ×napae House), and V. affinis (= V. ×subaffinis House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 6, p. 139. | FNA vol. 6, p. 140. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. arizonica, V. austiniae, V. cognata, V. crassula, V. galacifolia, V. lunellii, V. mccabeiana, V. nephrophylla var. arizonica, V. nephrophylla var. cognata, V. peramoena, V. pratincola, V. prionosepala, V. retusa, V. subjuncta, V. vagula | |
Name authority | Greene: Pittonia 3: 144. (1896) | Torrey & A. Gray: Fl. N. Amer. 1: 142. (1838) |
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