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kidney-leaf violet, Leconte violet, northern bog violet, violette néphrophylle

halberd-leaf or halberdleaf yellow violet, halberdleaf yellow violet

Habit Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age. Plants perennial, caulescent, not stoloniferous, 5–30 cm.
Stems

1(–2), erect, leafless proximally, leafy distally, glabrous or sparsely puberulent, from fleshy rhizome.

Leaves

basal, 4–7, ascending to erect;

stipules lanceolate, margins entire or fimbriate, apex acute;

petiole 2–25 cm, usually glabrous;

blade usually grayish green or purplish green abaxially, unlobed, ovate, reniform, or broadly reniform to orbiculate, 1–7 × 1–7 cm, somewhat fleshy, base broadly cordate or reniform to ± truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually glabrous, rarely sparsely pubescent.

basal and cauline;

basal: 0–1;

stipules ovate to lanceolate, margins laciniate with gland-tipped projections, apex usually acuminate;

petiole 3–11.5 cm, usually glabrous;

blade sometimes gray-green abaxially, usually mottled light green adaxially, widely or narrowly hastate to ovate, 2.5–4.4 × 1.8–3.2 cm, base hastate to cordate or truncate, margins serrate or crenate, ciliate or eciliate, apex acute, surfaces usually glabrous, sometimes scabrous;

cauline similar to basal except: leaves usually restricted to distal ends of naked stems;

petiole 1–2.5 cm;

blade 2–10 × 1.1–4.5 cm.

Peduncles

3–25 cm, usually glabrous.

1–5.3 cm, usually glabrous.

Flowers

sepals ovate, margins usually eciliate, auricles 1–2 mm;

petals deep bluish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, upper 2 sometimes sparsely bearded, lowest densely bearded or beardless, 10–28 mm, spur same color as petals, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers on erect to ascending peduncles.

sepals lanceolate to ovate, margins mostly eciliate, auricles 0.5–1 mm;

petals lemon-yellow usually on both surfaces, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded, lowest 9.5–12 mm, spur yellow to greenish, gibbous, 0.5–2 mm;

style head bearded; cleistogamous flowers axillary.

Capsules

ovoid, 5–10 mm, glabrous.

ovoid to ellipsoid, 6–8 mm, glabrous.

Seeds

beige to brown or dark brown, 1.5–2.5 mm.

beige to bronze, 2–2.5 mm.

2n

= 54.

= 12.

Viola nephrophylla

Viola hastata

Phenology Flowering Jan–Sep. Flowering late Mar–May.
Habitat Wet habitats in saturated soil in prairies, pastures, bogs, fens, sedge meadows, rocky shores of lakes and streams, limestone outcrops, gravelly calcareous stream beds Rich woods, chiefly mountains and piedmont
Elevation 100–3000 m (300–9800 ft) 50–2000 m (200–6600 ft)
Distribution
from FNA
AR; AZ; CA; CO; CT; IA; ID; IL; KS; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OH; OR; PA; SD; TX; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; GA; KY; MD; NC; NY; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola nephrophylla has the widest distribution in North America of any of the acaulescent blue violets.

There is some question whether the species we now know as Viola nephrophylla fits with the protologue and type specimen(s). McKinney examined the type designated by Greene; it appears that what most botanists have called V. nephrophylla may be an undescribed taxon. N. L. Gil-Ad (1997) believed Greene’s type contained plants that were genetically impure based on seed coat micromorphology and suggested the specimens might be a mixture of both orthospecies and introgressants.

Viola mccabeiana was described by M. S. Baker in 1940 from collections made in British Columbia. Later, Baker (1949b) discussed the reasons why the name V. mccabeiana needed to be abandoned. C. L. Hitchcock et al. (1955–1969, vol. 3) treated V. mccabeiana as synonymous with V. nephrophylla var. cognata. While Viola mccabeiana is recognized in Canada (L. Brouillet et al., http://data.canadensys.net/vascan/), we believe it is best included within V. nephrophylla.

Viola nephrophylla reportedly hybridizes with V. cucullata (= V. ×insessa House), V. sororia var. sororia (= V. ×napae House), and V. affinis (= V. ×subaffinis House).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

V. B. Baird (1942) commented that Viola hastata is more closely related to V. glabella than to any eastern species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 139. FNA vol. 6, p. 133.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. arizonica, V. austiniae, V. cognata, V. crassula, V. galacifolia, V. lunellii, V. mccabeiana, V. nephrophylla var. arizonica, V. nephrophylla var. cognata, V. peramoena, V. pratincola, V. prionosepala, V. retusa, V. subjuncta, V. vagula
Name authority Greene: Pittonia 3: 144. (1896) Michaux: Fl. Bor.-Amer. 2: 149. (1803)
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