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Macloskey's or northern white or small white or smooth white or wild white violet, Macloskey's violet, northern white violet, small white violet, smooth white violet, violette pâle

cream or cream-white or pale or striped or striped cream violet, creamy violet, striped cream violet, striped white violet, violette strièe

Habit Plants perennial, acaulescent, stoloniferous, 2–10 cm; stolons pale, often rooting and leafy at nodes; rhizome slender, fleshy. Plants perennial, caulescent, not stoloniferous, 10–60 cm.
Stems

1–4, ascending to erect (often declining during anthesis), glabrous or pubescent, on caudex from fleshy rhizome.

Leaves

basal, 2–6, ascending to erect;

stipules ovate to linear-lanceolate, margins entire or glandular-toothed, apex acute;

petiole 1–10 cm, strigose;

blade unlobed, reniform to ovate, 1–6.5 × 1–5.5 cm, base broadly or shallowly cordate, margins ± entire or shallowly crenate, eciliate, apex rounded to acute, surfaces usually glabrous, abaxial surfaces sometimes puberulent on proximal 1/2.

basal and cauline;

basal: 2–6;

stipules lanceolate to narrowly deltate, margins laciniate, apex acute;

petiole 3–6 cm, glabrous or puberulent;

blade ovate to reniform, 2–7 × 1–2.5 cm, base cordate, margins crenate to serrate, ciliate or eciliate, apex obtuse to acute, surfaces glabrous or pubescent;

cauline similar to basal except: stipules lanceolate, margins laciniate;

petiole 3–7 cm;

distal blades ovate to deltate, 1–6 × 1–4 cm, base cordate, apex acuminate to acute.

Peduncles

2.5–11(–21) cm, usually glabrous, sometimes villous.

5–12 cm, glabrous or pubescent.

Flowers

sepals lanceolate to ovate, margins eciliate, auricles 0.5–2 mm;

petals white on both surfaces, lower 3 purple-veined, lateral 2 bearded, rarely beardless, lowest 6–12 mm, spur white, gibbous, 1–2.5 mm;

style head beardless; cleistogamous flowers on ascending peduncles.

sepals lanceolate, margins ciliate or eciliate, auricles 2–3 mm;

petals white or cream on both surfaces, without yellow patch basally, lowest and usually lateral 2 purple-veined, lateral 2 and sometimes all densely bearded, lowest 10–18 mm, spur white, gibbous to elongated, 3–6 mm;

style head bearded; cleistogamous flowers axillary.

Capsules

ovoid, 5–9 mm, glabrous.

ellipsoid, 6–7 mm, glabrous.

Seeds

beige to bronze, 1–1.5 mm.

beige to bronze, 1.5–3 mm.

2n

= 24.

= 20.

Viola macloskeyi

Viola striata

Phenology Flowering Mar–Sep. Flowering Mar–Jun.
Habitat Bogs, wet meadows, seeps, lake margins, stream banks, floodplains, swampy woods, mesic roadside depressions, often among mosses Riparian or alluvial woods, floodplains in silty loam, meadows
Elevation 0–3600 m (0–11800 ft) 40–1000 m (100–3300 ft)
Distribution
from FNA
AZ; CA; CO; CT; DE; GA; IA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NV; NY; OH; OR; PA; RI; SC; SD; TN; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; SPM
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from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KY; MA; MD; MI; MO; NC; NH; NJ; NY; OH; OK; PA; SC; TN; VA; WI; WV; ON
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Discussion

Viola macloskeyi was described by Banks in 1824 as V. rotundifolia var. pallens, from specimens collected by Banks in Labrador. E. Brainerd made the combination V. pallens in 1905 and later (1924) detailed its troublesome nomenclatural history.

Viola macloskeyi was described in 1895 by F. E. Lloyd based on specimens collected at the base of Mount Hood, Oregon. M. S. Baker (1953) placed V. macloskeyi as a subspecies of V. pallens. He later (1953b) corrected that change based on priority and thus the taxon became known as V. macloskeyi subsp. pallens. N. H. Russell (1955) maintained that status, separating subspp. macloskeyi and pallens on the wider, spreading basal leaf lobes of subsp. pallens. McKinney, after examining specimens of both taxa, including types, concluded that the differences have been exaggerated and fall within the range of variation of a single species.

Viola macloskeyi occurs in small colonies; individual plants are interconnected by stolons.

In parts of its range, Viola macloskeyi often shares habitat with V. renifolia; it may be difficult to distinguish the two. The stolons of V. macloskeyi are useful for identification during summer months; V. renifolia plants lack stolons. Patterns of indument can be useful; V. renifolia is usually hairy and V. macloskeyi is usually glabrous (petioles sometimes pubescent).

Viola macloskeyi reportedly hybridizes with V. primulifolia var. primulifolia (= V. ×mollicula House).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dead, long-persistent stems of Viola striata are often present the following season. In flower, the plants are often mistaken for V. canadensis because the petals are whitish (H. E. Ballard 1992).

Viola striata reportedly hybridizes with V. labradorica (= V. ×eclipes H. E. Ballard), V. rostrata (= V. ×brauniae Grover ex Cooperrider), V. walteri var. appalachiensis (= V. ×wujekii H. E. Ballard), and V. walteri var. walteri (= V. ×cooperrideri H. E. Ballard).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 137. FNA vol. 6, p. 159.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. blanda subsp. macloskeyi, V. blanda var. macloskeyi, V. macloskeyi subsp. pallens, V. macloskeyi var. pallens, V. pallens, V. pallens subsp. macloskeyi, V. pallens var. subreptans, V. rotundifolia var. pallens Lophion striatum, V. conspersa var. masonii
Name authority F. E. Lloyd: Erythea 3: 74. (1895) Aiton: Hort. Kew. 3: 290. (1789)
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