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Macloskey's or northern white or small white or smooth white or wild white violet, Macloskey's violet, northern white violet, small white violet, smooth white violet, violette pâle

common dog-violet, dog violet, wood violet

Habit Plants perennial, acaulescent, stoloniferous, 2–10 cm; stolons pale, often rooting and leafy at nodes; rhizome slender, fleshy. Plants perennial, caulescent, not stoloniferous, 1.8–30 cm.
Stems

1–5, erect, ascending, or decumbent, glabrous or puberulent, on caudex from subligneous rhizome.

Leaves

basal, 2–6, ascending to erect;

stipules ovate to linear-lanceolate, margins entire or glandular-toothed, apex acute;

petiole 1–10 cm, strigose;

blade unlobed, reniform to ovate, 1–6.5 × 1–5.5 cm, base broadly or shallowly cordate, margins ± entire or shallowly crenate, eciliate, apex rounded to acute, surfaces usually glabrous, abaxial surfaces sometimes puberulent on proximal 1/2.

basal and cauline;

basal: 3–5;

stipules subulate or lanceolate, margins fimbriate, apex acute to acuminate;

petiole 2.5–12 cm, usually glabrous, rarely puberulent;

blade ovate to reniform, 2.3–5.5 × 2.6–5.3 cm, base cordate to deeply cordate, margins crenate, eciliate, apex acute or obtuse, abaxial surface glabrous or sparsely puberulent, adaxial surface usually sparsely puberulent, sometimes glabrous;

cauline similar to basal except: stipules ovate or lanceolate, margins fimbriate, apex long-acuminate;

petiole 1.3–6 cm;

blade ovate or broadly ovate, 1.8–5 × 1.5–4.2 cm, apex acute.

Peduncles

2.5–11(–21) cm, usually glabrous, sometimes villous.

2.8–10 cm, glabrous, sometimes puberulent.

Flowers

sepals lanceolate to ovate, margins eciliate, auricles 0.5–2 mm;

petals white on both surfaces, lower 3 purple-veined, lateral 2 bearded, rarely beardless, lowest 6–12 mm, spur white, gibbous, 1–2.5 mm;

style head beardless; cleistogamous flowers on ascending peduncles.

sepals lanceolate, margins eciliate;

auricles 1.8–2.5 mm (enlarged in fruit);

petals violet on both surfaces, lower 3 usually white basally, rarely violet, dark violet-veined, lateral 2 bearded, lowest 13–17 mm, spur white, rarely violet, elongated, 5–7 mm, tip straight or hooked, blunt;

style head beardless (surface papillose); cleistogamous flowers axillary.

Capsules

ovoid, 5–9 mm, glabrous.

ovoid or oblong, 8–12 mm, glabrous.

Seeds

beige to bronze, 1–1.5 mm.

pale to light brown, 1.8–2.1 mm.

2n

= 24.

= 40.

Viola macloskeyi

Viola riviniana

Phenology Flowering Mar–Sep. Flowering Mar–Sep.
Habitat Bogs, wet meadows, seeps, lake margins, stream banks, floodplains, swampy woods, mesic roadside depressions, often among mosses Roadsides and trails, sidewalk cracks, parks
Elevation 0–3600 m (0–11800 ft) 0–200 m (0–700 ft)
Distribution
from FNA
AZ; CA; CO; CT; DE; GA; IA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NV; NY; OH; OR; PA; RI; SC; SD; TN; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; SPM
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC; Eurasia; nw Africa [Introduced in North America; introduced in Australia]
Discussion

Viola macloskeyi was described by Banks in 1824 as V. rotundifolia var. pallens, from specimens collected by Banks in Labrador. E. Brainerd made the combination V. pallens in 1905 and later (1924) detailed its troublesome nomenclatural history.

Viola macloskeyi was described in 1895 by F. E. Lloyd based on specimens collected at the base of Mount Hood, Oregon. M. S. Baker (1953) placed V. macloskeyi as a subspecies of V. pallens. He later (1953b) corrected that change based on priority and thus the taxon became known as V. macloskeyi subsp. pallens. N. H. Russell (1955) maintained that status, separating subspp. macloskeyi and pallens on the wider, spreading basal leaf lobes of subsp. pallens. McKinney, after examining specimens of both taxa, including types, concluded that the differences have been exaggerated and fall within the range of variation of a single species.

Viola macloskeyi occurs in small colonies; individual plants are interconnected by stolons.

In parts of its range, Viola macloskeyi often shares habitat with V. renifolia; it may be difficult to distinguish the two. The stolons of V. macloskeyi are useful for identification during summer months; V. renifolia plants lack stolons. Patterns of indument can be useful; V. renifolia is usually hairy and V. macloskeyi is usually glabrous (petioles sometimes pubescent).

Viola macloskeyi reportedly hybridizes with V. primulifolia var. primulifolia (= V. ×mollicula House).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola riviniana has become established in several states on the Pacific Coast. It is cultivated and sold through nurseries in the United States. In the nursery trade in California and elsewhere, it is incorrectly referred to incorrectly as V. labradorica ‘Purpurea.’ In some plants shoots arise from adventitious buds on the roots (A. R. Clapham et al. 1987; T. Marcussen and T. Karlsson 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 137. FNA vol. 6, p. 154.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. blanda subsp. macloskeyi, V. blanda var. macloskeyi, V. macloskeyi subsp. pallens, V. macloskeyi var. pallens, V. pallens, V. pallens subsp. macloskeyi, V. pallens var. subreptans, V. rotundifolia var. pallens
Name authority F. E. Lloyd: Erythea 3: 74. (1895) Reichenbach: Iconogr. Bot. Pl. Crit. 1: 81, plate 95. (1823)
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