Viola macloskeyi
Viola quercetorum
Macloskey's or northern white or small white or smooth white or wild white violet, Macloskey's violet, northern white violet, small white violet, smooth white violet, violette pâle
goosefoot violet, goosefoot yellow violet, oakwoods violet
1–5, spreading to erect, leafy proximally and distally, usually elongated by end of season, puberulent to canescent, on caudex from subligneous rhizome.
basal, 2–6, ascending to erect;
stipules ovate to linear-lanceolate, margins entire or glandular-toothed, apex acute;
petiole 1–10 cm, strigose;
blade unlobed, reniform to ovate, 1–6.5 × 1–5.5 cm, base broadly or shallowly cordate, margins ± entire or shallowly crenate, eciliate, apex rounded to acute, surfaces usually glabrous, abaxial surfaces sometimes puberulent on proximal 1/2.
basal and cauline;
basal: 1–6;
stipules adnate to petiole, forming 2 linear, membranous wings, each wing with lanceolate to ± deltate projection, margins entire or laciniate, apex usually long-acuminate or divided into narrow, filiform processes;
petiole 1.9–9.5 cm, puberulent;
blade usually grayish green to whitish, sometimes ± purple-tinted abaxially, green to grayish green adaxially, ± ovate to ± orbiculate, deltate, pandurate, or broadly brownish purple abaxially, lower 3 dark brown-veined, lateral 2 sparsely to densely bearded, lowest 10–16 mm, spur yellow to reddish brown, gibbous, 1–2 mm;
style head bearded; cleistogamous flowers axillary.
2.5–11(–21) cm, usually glabrous, sometimes villous.
sepals lanceolate to ovate, margins eciliate, auricles 0.5–2 mm;
petals white on both surfaces, lower 3 purple-veined, lateral 2 bearded, rarely beardless, lowest 6–12 mm, spur white, gibbous, 1–2.5 mm;
style head beardless; cleistogamous flowers on ascending peduncles.
ovoid, 5–9 mm, glabrous.
ovoid to ellipsoid, 8–12 mm, puberulent.
beige to bronze, 1–1.5 mm.
medium brown, ± 2.7 mm.
= 24.
= 24.
Viola macloskeyi
Viola quercetorum
Viola macloskeyi was described by Banks in 1824 as V. rotundifolia var. pallens, from specimens collected by Banks in Labrador. E. Brainerd made the combination V. pallens in 1905 and later (1924) detailed its troublesome nomenclatural history.
Viola macloskeyi was described in 1895 by F. E. Lloyd based on specimens collected at the base of Mount Hood, Oregon. M. S. Baker (1953) placed V. macloskeyi as a subspecies of V. pallens. He later (1953b) corrected that change based on priority and thus the taxon became known as V. macloskeyi subsp. pallens. N. H. Russell (1955) maintained that status, separating subspp. macloskeyi and pallens on the wider, spreading basal leaf lobes of subsp. pallens. McKinney, after examining specimens of both taxa, including types, concluded that the differences have been exaggerated and fall within the range of variation of a single species.
Viola macloskeyi occurs in small colonies; individual plants are interconnected by stolons.
In parts of its range, Viola macloskeyi often shares habitat with V. renifolia; it may be difficult to distinguish the two. The stolons of V. macloskeyi are useful for identification during summer months; V. renifolia plants lack stolons. Patterns of indument can be useful; V. renifolia is usually hairy and V. macloskeyi is usually glabrous (petioles sometimes pubescent).
Viola macloskeyi reportedly hybridizes with V. primulifolia var. primulifolia (= V. ×mollicula House).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Herbarium specimens of Viola quercetorum and V. purpurea var. purpurea can be difficult to distinguish; pressing obliterates the undulate leaf margins of V. quercetorum. In Oregon, most collections of V. quercetorum are from yellow pine forests (A. Liston, pers. comm.).
Viola quercetorum hybridizes with V. douglasii (J. Clausen 1964).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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