Viola macloskeyi |
Viola pedata |
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Macloskey's or northern white or small white or smooth white or wild white violet, Macloskey's violet, northern white violet, small white violet, smooth white violet, violette pâle |
bird-foot violet |
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Habit | Plants perennial, acaulescent, stoloniferous, 2–10 cm; stolons pale, often rooting and leafy at nodes; rhizome slender, fleshy. | Plants perennial, acaulescent, not stoloniferous, 5–30 cm; rhizome thick, fleshy. | ||||
Leaves | basal, 2–6, ascending to erect; stipules ovate to linear-lanceolate, margins entire or glandular-toothed, apex acute; petiole 1–10 cm, strigose; blade unlobed, reniform to ovate, 1–6.5 × 1–5.5 cm, base broadly or shallowly cordate, margins ± entire or shallowly crenate, eciliate, apex rounded to acute, surfaces usually glabrous, abaxial surfaces sometimes puberulent on proximal 1/2. |
basal, 4–10, ascending to erect, deeply divided; stipules linear-lanceolate, margins entire, lacerate, or shallowly divided, apex acute; petiole 2–12 cm, glabrous or sparsely puberulent; blade 3–9(–10)-lobed, lobes similar in width and shape, spatulate, lanceolate ± linear, deltate, or ovate, 1–4 × 1–4 cm, base attenuate or broadly cordate to cuneate, margins entire, ciliate or eciliate, apex rounded to usually acute, surfaces usually glabrous, sometimes pubescent on abaxial veins. |
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Peduncles | 2.5–11(–21) cm, usually glabrous, sometimes villous. |
5–12 cm, glabrous or pubescent. |
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Flowers | sepals lanceolate to ovate, margins eciliate, auricles 0.5–2 mm; petals white on both surfaces, lower 3 purple-veined, lateral 2 bearded, rarely beardless, lowest 6–12 mm, spur white, gibbous, 1–2.5 mm; style head beardless; cleistogamous flowers on ascending peduncles. |
sepals lanceolate, margins mostly ciliate, at least proximally, auricles 1–2 mm; petals uniformly light to dark blue-violet on both surfaces or upper 2 darker adaxially, sometimes white, upper and lateral 2 often darker basally, lowest, seldom others, dark violet-veined, all beardless, lowest white basally, 12–24 mm, spur white, gibbous, 2–3 mm; style head beardless; cleistogamous flowers absent. |
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Capsules | ovoid, 5–9 mm, glabrous. |
ellipsoid, 6–10 mm, glabrous. |
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Seeds | beige to bronze, 1–1.5 mm. |
beige, mottled to brown, 1.4–3 mm. |
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2n | = 24. |
= 56. |
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Viola macloskeyi |
Viola pedata |
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Phenology | Flowering Mar–Sep. | |||||
Habitat | Bogs, wet meadows, seeps, lake margins, stream banks, floodplains, swampy woods, mesic roadside depressions, often among mosses | |||||
Elevation | 0–3600 m (0–11800 ft) | |||||
Distribution |
AZ; CA; CO; CT; DE; GA; IA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NV; NY; OH; OR; PA; RI; SC; SD; TN; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; SPM
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AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; ON
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Discussion | Viola macloskeyi was described by Banks in 1824 as V. rotundifolia var. pallens, from specimens collected by Banks in Labrador. E. Brainerd made the combination V. pallens in 1905 and later (1924) detailed its troublesome nomenclatural history. Viola macloskeyi was described in 1895 by F. E. Lloyd based on specimens collected at the base of Mount Hood, Oregon. M. S. Baker (1953) placed V. macloskeyi as a subspecies of V. pallens. He later (1953b) corrected that change based on priority and thus the taxon became known as V. macloskeyi subsp. pallens. N. H. Russell (1955) maintained that status, separating subspp. macloskeyi and pallens on the wider, spreading basal leaf lobes of subsp. pallens. McKinney, after examining specimens of both taxa, including types, concluded that the differences have been exaggerated and fall within the range of variation of a single species. Viola macloskeyi occurs in small colonies; individual plants are interconnected by stolons. In parts of its range, Viola macloskeyi often shares habitat with V. renifolia; it may be difficult to distinguish the two. The stolons of V. macloskeyi are useful for identification during summer months; V. renifolia plants lack stolons. Patterns of indument can be useful; V. renifolia is usually hairy and V. macloskeyi is usually glabrous (petioles sometimes pubescent). Viola macloskeyi reportedly hybridizes with V. primulifolia var. primulifolia (= V. ×mollicula House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 137. | FNA vol. 6, p. 143. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. blanda subsp. macloskeyi, V. blanda var. macloskeyi, V. macloskeyi subsp. pallens, V. macloskeyi var. pallens, V. pallens, V. pallens subsp. macloskeyi, V. pallens var. subreptans, V. rotundifolia var. pallens | |||||
Name authority | F. E. Lloyd: Erythea 3: 74. (1895) | Linnaeus: Sp. Pl. 2: 933. (1753) | ||||
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