Viola macloskeyi
Viola cuneata
Macloskey's or northern white or small white or smooth white or wild white violet, Macloskey's violet, northern white violet, small white violet, smooth white violet, violette pâle
northern two-eyed or wedge-leaf violet, northern two-eyed violet, wedge-leaf violet
1–3, usually erect, sometimes prostrate or ascending, glabrous, from shallow, fleshy rhizome or deep-seated caudex with fleshy roots.
basal, 2–6, ascending to erect;
stipules ovate to linear-lanceolate, margins entire or glandular-toothed, apex acute;
petiole 1–10 cm, strigose;
blade unlobed, reniform to ovate, 1–6.5 × 1–5.5 cm, base broadly or shallowly cordate, margins ± entire or shallowly crenate, eciliate, apex rounded to acute, surfaces usually glabrous, abaxial surfaces sometimes puberulent on proximal 1/2.
basal and cauline;
basal: 2–6;
stipules lanceolate, margins entire or gland-toothed, apex acute;
petiole 4.5–9.8 cm, glabrous;
blade purple-veined, orbiculate-ovate to deltate, 1–4 × 2.7–3.8 cm, usually shiny, leathery, base cuneate, margins serrate, eciliate, apex acute, mucronulate, surfaces glabrous;
cauline similar to basal except: stipules lanceolate to ovate, margins entire or erose to lacerate, with or without gland-tipped processes, apex acute to acuminate;
petiole 0.5–20 cm;
blade usually rhombic, sometimes orbiculate, 0.9–2.6 × 0.7–1.8 cm, margins crenate to ± serrate, apex acute to obtuse.
2.5–11(–21) cm, usually glabrous, sometimes villous.
1–10.5 cm, glabrous.
sepals lanceolate to ovate, margins eciliate, auricles 0.5–2 mm;
petals white on both surfaces, lower 3 purple-veined, lateral 2 bearded, rarely beardless, lowest 6–12 mm, spur white, gibbous, 1–2.5 mm;
style head beardless; cleistogamous flowers on ascending peduncles.
sepals lanceolate, margins eciliate, auricles 0.5–1 mm;
petals deep reddish violet abaxially, white adaxially, upper 2 sometimes with purple patch basally, lateral 2 with purple patch basally beyond smaller yellow area, usually bearded, sometimes beardless, lowest purple-veined with yellow area basally, 8–14 mm, spur yellow, gibbous, 1–2 mm;
style head bearded; cleistogamous flowers axillary.
ovoid, 5–9 mm, glabrous.
± spherical, 5–9 mm, glabrous.
beige to bronze, 1–1.5 mm.
deep brown-purple, 2.1–3 mm.
= 24.
Viola macloskeyi
Viola cuneata
Viola macloskeyi was described by Banks in 1824 as V. rotundifolia var. pallens, from specimens collected by Banks in Labrador. E. Brainerd made the combination V. pallens in 1905 and later (1924) detailed its troublesome nomenclatural history.
Viola macloskeyi was described in 1895 by F. E. Lloyd based on specimens collected at the base of Mount Hood, Oregon. M. S. Baker (1953) placed V. macloskeyi as a subspecies of V. pallens. He later (1953b) corrected that change based on priority and thus the taxon became known as V. macloskeyi subsp. pallens. N. H. Russell (1955) maintained that status, separating subspp. macloskeyi and pallens on the wider, spreading basal leaf lobes of subsp. pallens. McKinney, after examining specimens of both taxa, including types, concluded that the differences have been exaggerated and fall within the range of variation of a single species.
Viola macloskeyi occurs in small colonies; individual plants are interconnected by stolons.
In parts of its range, Viola macloskeyi often shares habitat with V. renifolia; it may be difficult to distinguish the two. The stolons of V. macloskeyi are useful for identification during summer months; V. renifolia plants lack stolons. Patterns of indument can be useful; V. renifolia is usually hairy and V. macloskeyi is usually glabrous (petioles sometimes pubescent).
Viola macloskeyi reportedly hybridizes with V. primulifolia var. primulifolia (= V. ×mollicula House).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Viola cuneata often occurs on serpentine-derived soil and is closely related to V. flettii, V. hallii, and V. ocellata (V. B. Baird 1942). Originally described as beardless by Watson, the lateral petals of flowers in the same population can be prominently bearded or essentially beardless. The cauline blades of Viola cuneata are often vertical, especially early in season.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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