Viola macloskeyi |
Viola canadensis |
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Macloskey's or northern white or small white or smooth white or wild white violet, Macloskey's violet, northern white violet, small white violet, smooth white violet, violette pâle |
Canada violet, Canada white violet, Canadian violet, Canadian white violet, rugose violet |
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Habit | Plants perennial, acaulescent, stoloniferous, 2–10 cm; stolons pale, often rooting and leafy at nodes; rhizome slender, fleshy. | Plants perennial, caulescent, not stoloniferous, 3–46(–60) cm, with branching rhizomes forming colonies or not. | ||||||||
Stems | 1–3(4), usually erect to ascending, glabrous or puberulent, from fleshy or subligneous rhizome. |
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Leaves | basal, 2–6, ascending to erect; stipules ovate to linear-lanceolate, margins entire or glandular-toothed, apex acute; petiole 1–10 cm, strigose; blade unlobed, reniform to ovate, 1–6.5 × 1–5.5 cm, base broadly or shallowly cordate, margins ± entire or shallowly crenate, eciliate, apex rounded to acute, surfaces usually glabrous, abaxial surfaces sometimes puberulent on proximal 1/2. |
basal and cauline; basal: 1–5; stipules ± oblong, ovate, or lanceolate, margins entire, sometimes glandular, apex acuminate to cuspidate; petiole 1.1–23 cm, glabrous or puberulent; blade ovate to broadly ovate or ovate-reniform, 0.7–12.4 × 0.9–11.1(–12.3) cm, base cordate, subcordate, or truncate, margins crenate, crenulate, or serrulate, ciliate (sometimes only on proximal 1/2) or eciliate, apex acute to acuminate, surfaces glabrous or puberulent (often only on veins); cauline similar to basal except: stipules also deltate, margins also erose or laciniate, apex acute, long-acuminate to cuspidate, or ± truncate, occasionally 2- or 3-fid; petiole 0.1–6.9(–15.2) cm; blade ovate to deltate, 1.2–7.7 × 0.8–7.8 cm, base cordate to truncate, margins crenate or crenulate to ± serrulate. |
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Peduncles | 2.5–11(–21) cm, usually glabrous, sometimes villous. |
1–6.1 cm, glabrate to puberulent, sometimes glabrous below bracteoles. |
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Flowers | sepals lanceolate to ovate, margins eciliate, auricles 0.5–2 mm; petals white on both surfaces, lower 3 purple-veined, lateral 2 bearded, rarely beardless, lowest 6–12 mm, spur white, gibbous, 1–2.5 mm; style head beardless; cleistogamous flowers on ascending peduncles. |
sepals lanceolate, margins usually eciliate, auricles 0.5–1.3 mm; petals white adaxially, upper 2 and lower 3 tinged soft reddish violet abaxially, rarely white on both surfaces, all petals usually with yellow patch basally, lower 3 usually purple-veined, lateral 2 bearded, lowest 5.5–20 mm, spur white, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary or absent. |
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Capsules | ovoid, 5–9 mm, glabrous. |
ovoid to ellipsoid, 2.5–10 mm, sometimes muriculate, glabrous or puberulent. |
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Seeds | beige to bronze, 1–1.5 mm. |
brown to dark brown or purplish black, 1.5–2.5 mm. |
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2n | = 24. |
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Viola macloskeyi |
Viola canadensis |
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Phenology | Flowering Mar–Sep. | |||||||||
Habitat | Bogs, wet meadows, seeps, lake margins, stream banks, floodplains, swampy woods, mesic roadside depressions, often among mosses | |||||||||
Elevation | 0–3600 m (0–11800 ft) | |||||||||
Distribution |
AZ; CA; CO; CT; DE; GA; IA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; ND; NH; NJ; NV; NY; OH; OR; PA; RI; SC; SD; TN; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; SPM
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AK; AL; AR; AZ; CO; CT; GA; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OR; PA; RI; SC; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; NT; ON; QC; SK; YT
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Discussion | Viola macloskeyi was described by Banks in 1824 as V. rotundifolia var. pallens, from specimens collected by Banks in Labrador. E. Brainerd made the combination V. pallens in 1905 and later (1924) detailed its troublesome nomenclatural history. Viola macloskeyi was described in 1895 by F. E. Lloyd based on specimens collected at the base of Mount Hood, Oregon. M. S. Baker (1953) placed V. macloskeyi as a subspecies of V. pallens. He later (1953b) corrected that change based on priority and thus the taxon became known as V. macloskeyi subsp. pallens. N. H. Russell (1955) maintained that status, separating subspp. macloskeyi and pallens on the wider, spreading basal leaf lobes of subsp. pallens. McKinney, after examining specimens of both taxa, including types, concluded that the differences have been exaggerated and fall within the range of variation of a single species. Viola macloskeyi occurs in small colonies; individual plants are interconnected by stolons. In parts of its range, Viola macloskeyi often shares habitat with V. renifolia; it may be difficult to distinguish the two. The stolons of V. macloskeyi are useful for identification during summer months; V. renifolia plants lack stolons. Patterns of indument can be useful; V. renifolia is usually hairy and V. macloskeyi is usually glabrous (petioles sometimes pubescent). Viola macloskeyi reportedly hybridizes with V. primulifolia var. primulifolia (= V. ×mollicula House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 137. | FNA vol. 6, p. 125. | ||||||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||||||
Sibling taxa | ||||||||||
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Synonyms | V. blanda subsp. macloskeyi, V. blanda var. macloskeyi, V. macloskeyi subsp. pallens, V. macloskeyi var. pallens, V. pallens, V. pallens subsp. macloskeyi, V. pallens var. subreptans, V. rotundifolia var. pallens | |||||||||
Name authority | F. E. Lloyd: Erythea 3: 74. (1895) | Linnaeus: Sp. Pl. 2: 936. (1753) | ||||||||
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