Viola lobata |
Viola subsinuata |
|||||
---|---|---|---|---|---|---|
moose horn violet, pine violet, yellow wood violet |
early blue violet, wavy-leaf violet, wood violet |
|||||
Habit | Plants perennial, caulescent, not stoloniferous, 5–46 cm. | Plants perennial, acaulescent, not stoloniferous, 10–30 cm; rhizome thick, fleshy. | ||||
Stems | 1–3, erect, leafless proximally, leafy distally, glabrous or puberulent, from subligneous rhizome. |
|||||
Leaves | basal and cauline; basal: 0–2, unlobed or palmately 3–11-lobed; stipules ovate to lanceolate, margins ± entire or serrate, apex acute; petiole 5–27 cm, glabrous or puberulent; blade sometimes glaucous, deltate to reniform, 3.5–8.5 × 4.5–13.5 cm, base cordate, truncate, or attenuate, margins on unlobed leaves coarsely dentate-serrate, margins on lobed leaves usually entire, sometimes few-toothed, ciliate (sometimes only proximal 1/2) or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: distal on naked stems, unlobed and divided leaves can occur on same plant, if divided, palmately 3–12-lobed; stipules sometimes large and ± leaflike, margins entire, lacerate, or laciniate, sometimes with gland-tipped projections, apex acute to acuminate; petiole 0.2–8.8 cm; blade reniform to reniform-cordate, ± ovate, deltate, or rhombic, 1.5–5.5 × 1.4–10 cm, base cordate, subcordate, truncate, or attenuate, margins entire, crenate-serrate, or dentate, or coarsely lacerate to deeply serrate, often entire distally, ciliate or rarely eciliate, apex acute to obtuse, often long-tapered, mucronulate. |
basal, 2–11, ascending to erect; stipules linear-lanceolate, margins entire, apex acute; petiole 5–25 cm, glabrous or pubescent; mid-season leaf blades incised or lobed throughout, earliest leaf blades lobed (plants homophyllous), similar to mid-season blades, blade 5–9(–16)-lobed, sinuses usually narrower, shallower toward leaf base, middle and lateral blade lobes differ in width and/or shape, middle lobes narrowly deltate to narrowly elliptic, lateral lobes narrowly elliptic, lanceolate, or falcate, 1–11 × 1–12 cm, base truncate to cordate, margins entire or crenate, ciliate or eciliate, apex acute, mucronulate, surfaces usually pubescent throughout or on veins. |
||||
Peduncles | 2–13 cm, glabrous or pubescent. |
5–15 cm, glabrous or pubescent. |
||||
Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 0.1–1 mm; petals deep lemon-yellow adaxially, usually upper 2 and sometimes lateral 2 brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined basally, lateral 2 bearded, lowest 8–19 mm, spur yellow to greenish, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to dark blue-violet on both surfaces, lower 3 and upper 2 sometimes purple-veined, lateral 2 bearded, lowest sometimes bearded, 15–25 mm, spur color same as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
||||
Capsules | ellipsoid-ovoid, 6–16 mm, glabrous. |
ellipsoid, 8–12 mm, glabrous. |
||||
Seeds | light brown, blotched or streaked with brown, shiny, 2.1–2.7 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
||||
2n | = 12. |
= 54. |
||||
Viola lobata |
Viola subsinuata |
|||||
Phenology | Flowering Apr–Jun. | |||||
Habitat | Rich woods | |||||
Elevation | 100–3000 m (300–9800 ft) | |||||
Distribution |
CA; OR; nw Mexico
|
CT; DC; DE; GA; IL; KY; MA; MD; MI; NC; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; ON
|
||||
Discussion | Varieties 2 (2 in the flora). V. B. Baird (1942) noted that Viola lobata was more closely related to V. tripartita than to any western Viola. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
L. E. McKinney (1992) described the misconceptions surrounding Viola subsinuata that many came to think of as the heterophyllous V. palmata, whereas V. subsinuata is homophyllous. As E. Brainerd (1910, 1921) pointed out, heterophylly versus homophylly is an important and steadfast character difference in acaulescent Viola species. N. L. Gil-Ad (1997) chose not to recognize V. subsinuata, suggesting that it represented a hybrid or introgressant. H. E. Ballard (2000) and A. Haines (2011) both recognized V. subsinuata as a species. Viola subsinuata is a state historical species in Rhode Island, where it was last documented in 1941 (R. W. Enser, http://rinhs.org/wp-content/uploads/ri_rare_plants_2007.pdf). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||
Key |
|
|||||
Source | FNA vol. 6, p. 137. | FNA vol. 6, p. 159. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. emarginata var. subsinuata, V. palmata var. angelliae | |||||
Name authority | Bentham: Pl. Hartw., 298. (1849) | (Greene) Greene: Pittonia 4: 4. (1899) | ||||
Web links |