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moose horn violet, pine violet, yellow wood violet

bog white violet, Howell's violet, lance-leaf or bog white violet, lance-leaf violet, strap-leaf violet, violette lancéolée, white bog violet

Habit Plants perennial, caulescent, not stoloniferous, 5–46 cm. Plants perennial, acaulescent, stoloniferous, 5–30 cm; stolons pale, often rooting and leafy at nodes; rhizome slender, fleshy.
Stems

1–3, erect, leafless proximally, leafy distally, glabrous or puberulent, from subligneous rhizome.

Leaves

basal and cauline;

basal: 0–2, unlobed or palmately 3–11-lobed;

stipules ovate to lanceolate, margins ± entire or serrate, apex acute;

petiole 5–27 cm, glabrous or puberulent;

blade sometimes glaucous, deltate to reniform, 3.5–8.5 × 4.5–13.5 cm, base cordate, truncate, or attenuate, margins on unlobed leaves coarsely dentate-serrate, margins on lobed leaves usually entire, sometimes few-toothed, ciliate (sometimes only proximal 1/2) or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;

cauline similar to basal except: distal on naked stems, unlobed and divided leaves can occur on same plant, if divided, palmately 3–12-lobed;

stipules sometimes large and ± leaflike, margins entire, lacerate, or laciniate, sometimes with gland-tipped projections, apex acute to acuminate;

petiole 0.2–8.8 cm;

blade reniform to reniform-cordate, ± ovate, deltate, or rhombic, 1.5–5.5 × 1.4–10 cm, base cordate, subcordate, truncate, or attenuate, margins entire, crenate-serrate, or dentate, or coarsely lacerate to deeply serrate, often entire distally, ciliate or rarely eciliate, apex acute to obtuse, often long-tapered, mucronulate.

basal, 2–6(–9), ascending to erect;

stipules linear-lanceolate, margins entire or irregularly lacerate (at least distally), apex acute;

petiole 2–12 cm, glabrous or pubescent;

blade unlobed, lanceolate or narrowly elliptic to nearly linear, 2.5–12 × 0.7–2.5 cm, longer than wide, base attenuate, margins serrate, mostly eciliate, apex acute, mucronulate, surfaces glabrous.

Peduncles

2–13 cm, glabrous or pubescent.

2–17 cm, usually pubescent.

Flowers

sepals lanceolate, margins ciliate or eciliate, auricles 0.1–1 mm;

petals deep lemon-yellow adaxially, usually upper 2 and sometimes lateral 2 brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined basally, lateral 2 bearded, lowest 8–19 mm, spur yellow to greenish, gibbous, 0.5–2 mm;

style head bearded; cleistogamous flowers axillary.

sepals ovate to lanceolate, margins mostly eciliate, auricles 1–2 mm;

petals white on both surfaces, lowest and sometimes lateral 2 purple-veined, lateral 2 sparsely bearded or beardless, lowest 7–12 mm, spur white, gibbous, 1–2 mm;

style head beardless; cleistogamous flowers on prostrate to ascending peduncles.

Capsules

ellipsoid-ovoid, 6–16 mm, glabrous.

ellipsoid, 5–8 mm, glabrous.

Seeds

light brown, blotched or streaked with brown, shiny, 2.1–2.7 mm.

beige to bronze, 1.5–2.5 mm.

2n

= 12.

= 24.

Viola lobata

Viola lanceolata

Phenology Flowering Mar–May.
Habitat Open to semi-open wet areas, bogs, meadows, pond and lake shores, stream banks, seasonally inundated depressions
Elevation 0–2000 m (0–6600 ft)
Distribution
from FNA
CA; OR; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; OR; PA; RI; SC; TN; TX; VA; VT; WA; WI; WV; BC; NB; NF; NS; ON; PE; QC; SPM
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

V. B. Baird (1942) noted that Viola lobata was more closely related to V. tripartita than to any western Viola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola lanceolata occurs in small colonies; individual plants are interconnected by stolons.

Viola lanceolata subsp. vittata was recognized by N. H. Russell (1955) based on its linear leaf blade shape. The range in leaf blade shape appears to have no distinct line of demarcation. Some believe that leaf shape differences and near restriction to the coastal plain support recognition at some level. Studies are necessary to resolve this issue.

J. H. Shultze (1946) reported that Viola lanceolata was introduced into Washington in the early 1900s primarily from Cape Cod and Wisconsin as a result of importation of cranberry vines. In British Columbia, it is known from Lulu Island, where it was introduced from eastern North America (G. W. Douglas et al. 1998–2002, vol. 5).

Viola lanceolata reportedly hybridizes with V. primulifolia var. primulifolia (= V. ×modesta House) and V. macloskeyi (= V. ×sublanceolata House).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cauline leaf blades usually deeply divided, seldom unlobed, reniform, ± ovate, or deltate, if divided, palmately 3–12-lobed, lobe margins usually entire.
var. lobata
1. Cauline leaf blades unlobed, deltate to rhombic, ovate, or reniform-cordate, margins ± serrate, crenate-serrate, or dentate to deeply serrate or coarsely lacerate, often entire distally.
var. integrifolia
Source FNA vol. 6, p. 137. FNA vol. 6, p. 135.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. lobata var. integrifolia, V. lobata var. lobata
Synonyms V. lanceolata subsp. vittata, V. lanceolata var. vittata, V. vittata
Name authority Bentham: Pl. Hartw., 298. (1849) Linnaeus: Sp. Pl. 2: 934. (1753)
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