Viola lobata |
Viola labradorica |
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moose horn violet, pine violet, yellow wood violet |
alpine violet, American dog or Labrador or alpine violet, American dog violet, dog violet, Labrador violet, violette du Labrador |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–46 cm. | Plants perennial, caulescent, not stoloniferous, 5–20 cm. | ||||
Stems | 1–3, erect, leafless proximally, leafy distally, glabrous or puberulent, from subligneous rhizome. |
1–5, erect (later reclining), glabrous, on caudex from subligneous rhizome. |
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Leaves | basal and cauline; basal: 0–2, unlobed or palmately 3–11-lobed; stipules ovate to lanceolate, margins ± entire or serrate, apex acute; petiole 5–27 cm, glabrous or puberulent; blade sometimes glaucous, deltate to reniform, 3.5–8.5 × 4.5–13.5 cm, base cordate, truncate, or attenuate, margins on unlobed leaves coarsely dentate-serrate, margins on lobed leaves usually entire, sometimes few-toothed, ciliate (sometimes only proximal 1/2) or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: distal on naked stems, unlobed and divided leaves can occur on same plant, if divided, palmately 3–12-lobed; stipules sometimes large and ± leaflike, margins entire, lacerate, or laciniate, sometimes with gland-tipped projections, apex acute to acuminate; petiole 0.2–8.8 cm; blade reniform to reniform-cordate, ± ovate, deltate, or rhombic, 1.5–5.5 × 1.4–10 cm, base cordate, subcordate, truncate, or attenuate, margins entire, crenate-serrate, or dentate, or coarsely lacerate to deeply serrate, often entire distally, ciliate or rarely eciliate, apex acute to obtuse, often long-tapered, mucronulate. |
basal and cauline; basal: 1–5; stipules mostly lanceolate-linear, margins usually laciniate with gland-tipped projections, apex acute; petiole 1–9.5 cm, glabrous or puberulent; blade reniform or ovate to deltate, 1.4–5.5 × 1.8–5.3 cm, base cordate, not decurrent on petiole, margins crenate or serrate, mostly eciliate, apex acute to obtuse, surfaces usually glabrous, sometimes pubescent adaxially near margin; cauline similar to basal except: stipules lanceolate to ovate, margins ± entire or laciniate, projections often gland-tipped, apex acuminate; petiole 0.5–3.6 cm, glabrous; blade narrowly ovate to orbiculate, 1.8–2.9 × 1.5–3.1 cm, base deeply cordate to truncate, margins regularly crenate, apex obtuse to broadly rounded or apiculate. |
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Peduncles | 2–13 cm, glabrous or pubescent. |
5–9 cm, glabrous or pubescent. |
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Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 0.1–1 mm; petals deep lemon-yellow adaxially, usually upper 2 and sometimes lateral 2 brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined basally, lateral 2 bearded, lowest 8–19 mm, spur yellow to greenish, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins ciliate or eciliate, auricles not enlarged in fruit, 1–1.5 mm; petals usually lavender-violet to violet on both surfaces, rarely white, lower 3 white basally, darker violet-veined, lateral 2 sparsely bearded, lowest 9–16 mm, spur white to pinkish violet, elongated, 4–8 mm, tip usually straight, sometimes curved up; style head beardless; cleistogamous flowers unknown. |
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Capsules | ellipsoid-ovoid, 6–16 mm, glabrous. |
ovoid to ellipsoid, 5–7 mm, glabrous. |
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Seeds | light brown, blotched or streaked with brown, shiny, 2.1–2.7 mm. |
beige to bronze, 1.5–2 mm. |
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2n | = 12. |
= 20. |
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Viola lobata |
Viola labradorica |
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Phenology | Flowering Apr–Aug. | |||||
Habitat | Rich, mesic to wet woods, swamps, bogs | |||||
Elevation | 50–3000 m (200–9800 ft) | |||||
Distribution |
CA; OR; nw Mexico
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AL; CO; CT; DC; DE; GA; IL; IN; KY; MA; MD; ME; MI; MN; NC; ND; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; MB; NB; NL; NS; NT; ON; PE; QC; SK; SPM; Greenland
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Discussion | Varieties 2 (2 in the flora). V. B. Baird (1942) noted that Viola lobata was more closely related to V. tripartita than to any western Viola. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
H. E. Ballard (1994) made a compelling case for including Viola conspersa within V. labradorica, which is followed here. Ballard (1992) did not recognize V. labradorica as occurring in Colorado. Several western herbaria have collections identified as V. labradorica (ASC, CS, KHD, RM, RMBL; http://swbiodiversity.org/seinet/index.pho). Labels on some collections at KHD imply that V. labradorica is considered synonymous with V. adunca var. bellidifolia. Viola labradorica reportedly hybridizes with V. adunca, V. rostrata (= V. ×malteana House), and V. striata (= V. ×eclipes H. E. Ballard), and less frequently with V. walteri var. appalachiensis and var. walteri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 137. | FNA vol. 6, p. 135. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. adunca var. minor, V. canina var. muhlenbergii, V. conspersa, V. leucopetala, V. muhlenbergiana, V. muhlenbergiana var. minor, V. muhlenbergii | |||||
Name authority | Bentham: Pl. Hartw., 298. (1849) | Schrank: Denkschr. Kõnigl.-Baier. Bot. Ges. Regensburg 1(2): 12. (1818) | ||||
Web links |