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moose horn violet, pine violet, yellow wood violet

southern woodland violet

Habit Plants perennial, caulescent, not stoloniferous, 5–46 cm. Plants perennial, acaulescent, not stoloniferous, 2–15 cm; rhizome thick, fleshy.
Stems

1–3, erect, leafless proximally, leafy distally, glabrous or puberulent, from subligneous rhizome.

Leaves

basal and cauline;

basal: 0–2, unlobed or palmately 3–11-lobed;

stipules ovate to lanceolate, margins ± entire or serrate, apex acute;

petiole 5–27 cm, glabrous or puberulent;

blade sometimes glaucous, deltate to reniform, 3.5–8.5 × 4.5–13.5 cm, base cordate, truncate, or attenuate, margins on unlobed leaves coarsely dentate-serrate, margins on lobed leaves usually entire, sometimes few-toothed, ciliate (sometimes only proximal 1/2) or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;

cauline similar to basal except: distal on naked stems, unlobed and divided leaves can occur on same plant, if divided, palmately 3–12-lobed;

stipules sometimes large and ± leaflike, margins entire, lacerate, or laciniate, sometimes with gland-tipped projections, apex acute to acuminate;

petiole 0.2–8.8 cm;

blade reniform to reniform-cordate, ± ovate, deltate, or rhombic, 1.5–5.5 × 1.4–10 cm, base cordate, subcordate, truncate, or attenuate, margins entire, crenate-serrate, or dentate, or coarsely lacerate to deeply serrate, often entire distally, ciliate or rarely eciliate, apex acute to obtuse, often long-tapered, mucronulate.

basal, 2–4, prostrate or nearly so;

stipules linear-lanceolate, margins entire, apex acute;

petiole 1–10 cm, usually glabrous;

blade unlobed, reniform to ovate, 1–6 × 1–5 cm, base cordate, margins serrate, ciliate or eciliate, apex rounded, adaxial leaf surface with silvery strigose patches.

Peduncles

2–13 cm, glabrous or pubescent.

2–12 cm, glabrous.

Flowers

sepals lanceolate, margins ciliate or eciliate, auricles 0.1–1 mm;

petals deep lemon-yellow adaxially, usually upper 2 and sometimes lateral 2 brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined basally, lateral 2 bearded, lowest 8–19 mm, spur yellow to greenish, gibbous, 0.5–2 mm;

style head bearded; cleistogamous flowers axillary.

sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm;

petals light to deep reddish violet to blue-violet on both surfaces, 3 lower whitish basally, dark violet-veined, lateral 2 densely bearded, lowest 11–17 mm, spur violet, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers on prostrate to ascending peduncles.

Capsules

ellipsoid-ovoid, 6–16 mm, glabrous.

ellipsoid, 8–12 mm, glabrous.

Seeds

light brown, blotched or streaked with brown, shiny, 2.1–2.7 mm.

beige, mottled to bronze, 1–1.5 mm.

2n

= 12.

= 54.

Viola lobata

Viola hirsutula

Phenology Flowering Mar–Jun.
Habitat Dry to mesic deciduous woodlands
Elevation 100–1000 m (300–3300 ft)
Distribution
from FNA
CA; OR; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DC; DE; FL; GA; IN; KY; MD; MS; NC; NJ; NY; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

V. B. Baird (1942) noted that Viola lobata was more closely related to V. tripartita than to any western Viola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola hirsutula exhibits little phenotypic plasticity. Although considered by N. L. Gil-Ad (1997) to be a putative hybrid, he did not identify the potential parents and chose not to recognize it. A. Cronquist in H. A. Gleason and Cronquist (1991) considered V. hirsutula a form of V. villosa. Viola hirsutula is considered one of the more distinct species throughout its range by McKinney because of the pattern of silvery strigose patches on adaxial leaf surfaces and a more prostrate growth habit than similar species. He believes V. hirsutula is one of the least variable species in the flora. Viola hirsutula appears to occur much less frequently than herbarium collections suggest. Some herbarium specimens are misidentified and represent depauperate plants, usually of V. sororia. These specimens are often hairy on the adaxial leaf surfaces but lack the silvery patches.

Viola hirsutula reportedly hybridizes with V. sororia var. sororia (= V. ×cordifolia [Nuttall] Schweinitz) and V. sagittata var. sagittata (= V. ×redacta House).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cauline leaf blades usually deeply divided, seldom unlobed, reniform, ± ovate, or deltate, if divided, palmately 3–12-lobed, lobe margins usually entire.
var. lobata
1. Cauline leaf blades unlobed, deltate to rhombic, ovate, or reniform-cordate, margins ± serrate, crenate-serrate, or dentate to deeply serrate or coarsely lacerate, often entire distally.
var. integrifolia
Source FNA vol. 6, p. 137. FNA vol. 6, p. 133.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. lobata var. integrifolia, V. lobata var. lobata
Name authority Bentham: Pl. Hartw., 298. (1849) Brainerd: Rhodora 9: 98. (1907)
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