Viola lobata |
Viola guadalupensis |
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moose horn violet, pine violet, yellow wood violet |
Guadalupe Mountains violet, Guadalupe violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–46 cm. | Plants perennial, caulescent, not stoloniferous, 1–10 cm. | ||||
Stems | 1–3, erect, leafless proximally, leafy distally, glabrous or puberulent, from subligneous rhizome. |
1–5, decumbent to erect, leafy proximally and distally, glabrous, on caudex from fleshy rhizome. |
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Leaves | basal and cauline; basal: 0–2, unlobed or palmately 3–11-lobed; stipules ovate to lanceolate, margins ± entire or serrate, apex acute; petiole 5–27 cm, glabrous or puberulent; blade sometimes glaucous, deltate to reniform, 3.5–8.5 × 4.5–13.5 cm, base cordate, truncate, or attenuate, margins on unlobed leaves coarsely dentate-serrate, margins on lobed leaves usually entire, sometimes few-toothed, ciliate (sometimes only proximal 1/2) or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: distal on naked stems, unlobed and divided leaves can occur on same plant, if divided, palmately 3–12-lobed; stipules sometimes large and ± leaflike, margins entire, lacerate, or laciniate, sometimes with gland-tipped projections, apex acute to acuminate; petiole 0.2–8.8 cm; blade reniform to reniform-cordate, ± ovate, deltate, or rhombic, 1.5–5.5 × 1.4–10 cm, base cordate, subcordate, truncate, or attenuate, margins entire, crenate-serrate, or dentate, or coarsely lacerate to deeply serrate, often entire distally, ciliate or rarely eciliate, apex acute to obtuse, often long-tapered, mucronulate. |
cauline; stipules lanceolate to ovate- or oblong-lanceolate or linear, margins sparingly glandular-fimbriate, apex acute; petiole 2–6 cm, glabrous; blade ovate to ovate-deltate or ovate-lanceolate, 1.2–2.4 × 0.7–1.3 cm, base broadly cuneate to rounded or truncate, margins entire or with 1–3 crenations on proximal 1/2, eciliate, apex acute to rounded, surfaces glabrous, sometimes with a few short hairs on veins abaxially. |
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Peduncles | 2–13 cm, glabrous or pubescent. |
3.5–6 cm, glabrous. |
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Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 0.1–1 mm; petals deep lemon-yellow adaxially, usually upper 2 and sometimes lateral 2 brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined basally, lateral 2 bearded, lowest 8–19 mm, spur yellow to greenish, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals linear to linear-lanceolate, margins eciliate, auricles 0.5–1.5 mm; petals deep lemon-yellow adaxially, upper 2 reddish brown abaxially, lateral 2 and lowest dark brown-veined basally, lateral 2 bearded, lowest 7–11 mm, spur yellow, gibbous, 1–1.4 mm; style head bearded; cleistogamous flowers absent. |
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Capsules | ellipsoid-ovoid, 6–16 mm, glabrous. |
ovoid, 3–4.5 mm, glabrous. |
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Seeds | light brown, blotched or streaked with brown, shiny, 2.1–2.7 mm. |
light brown, ± 2 mm. |
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2n | = 12. |
= 24. |
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Viola lobata |
Viola guadalupensis |
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Phenology | Flowering May. | |||||
Habitat | Openings and narrow ledges on limestone rock faces | |||||
Elevation | 2600 m (8500 ft) | |||||
Distribution |
CA; OR; nw Mexico
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TX |
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Discussion | Varieties 2 (2 in the flora). V. B. Baird (1942) noted that Viola lobata was more closely related to V. tripartita than to any western Viola. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Viola guadalupensis is known only from the eastern rim of the Guadalupe Mountains in Culberson County. Powell and Wauer noted that it is the only yellow-flowered violet known in the Guadalupe Mountains and appears to be related to V. nuttallii and V. vallicola. K. W. Allred (2008) stated that a report of this species in New Mexico by J. T. Kartesz and C. A. Meacham (1999) requires verification. K. Haskins (pers. comm.) reported that experiments are currently being conducted to propagate plants of V. guadalupensis via cell tissue culture. Chloroplast (trnL-F spacer) and low-copy nuclear gene (GPI) phylogenies indicate that Viola guadalupensis is an alloploid that originated through hybridization between an unidentified member of subsect. Canadenses (the paternal parent) and a member of the V. nuttallii complex (the maternal parent), of sect. Chamaemelanium (T. Marcussen et al. 2011). Evidence reported by these authors from a fossil-calibrated relaxed clock dating analysis showed the estimated maximum age of V. guadalupensis to be (5.7–)8.6(–11.6) million years. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 137. | FNA vol. 6, p. 132. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Name authority | Bentham: Pl. Hartw., 298. (1849) | A. M. Powell & Wauer: Sida 14: 1, fig. 1. (1990) | ||||
Web links |