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moose horn violet, pine violet, yellow wood violet

Clausen's violet

Habit Plants perennial, caulescent, not stoloniferous, 5–46 cm. Plants perennial, acaulescent, not stoloniferous, 3–20 cm; rhizome short, slender, fleshy.
Stems

1–3, erect, leafless proximally, leafy distally, glabrous or puberulent, from subligneous rhizome.

Leaves

basal and cauline;

basal: 0–2, unlobed or palmately 3–11-lobed;

stipules ovate to lanceolate, margins ± entire or serrate, apex acute;

petiole 5–27 cm, glabrous or puberulent;

blade sometimes glaucous, deltate to reniform, 3.5–8.5 × 4.5–13.5 cm, base cordate, truncate, or attenuate, margins on unlobed leaves coarsely dentate-serrate, margins on lobed leaves usually entire, sometimes few-toothed, ciliate (sometimes only proximal 1/2) or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;

cauline similar to basal except: distal on naked stems, unlobed and divided leaves can occur on same plant, if divided, palmately 3–12-lobed;

stipules sometimes large and ± leaflike, margins entire, lacerate, or laciniate, sometimes with gland-tipped projections, apex acute to acuminate;

petiole 0.2–8.8 cm;

blade reniform to reniform-cordate, ± ovate, deltate, or rhombic, 1.5–5.5 × 1.4–10 cm, base cordate, subcordate, truncate, or attenuate, margins entire, crenate-serrate, or dentate, or coarsely lacerate to deeply serrate, often entire distally, ciliate or rarely eciliate, apex acute to obtuse, often long-tapered, mucronulate.

basal, 2–10, often prostrate, sometimes ascending;

stipules narrowly lanceolate, margins faintly glandular-toothed, apex acute or obtuse;

petiole 5–11 cm, glabrous;

blade unlobed, ± deltate, 3–5 × 4–5 cm, base ± truncate, margins serrate, ciliate or eciliate, apex obtuse, surfaces usually glabrous, sometimes sparsely pubescent abaxially.

Peduncles

2–13 cm, glabrous or pubescent.

8–14 cm, glabrous.

Flowers

sepals lanceolate, margins ciliate or eciliate, auricles 0.1–1 mm;

petals deep lemon-yellow adaxially, usually upper 2 and sometimes lateral 2 brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined basally, lateral 2 bearded, lowest 8–19 mm, spur yellow to greenish, gibbous, 0.5–2 mm;

style head bearded; cleistogamous flowers axillary.

sepals lanceolate to ovate, margins eciliate or ciliate around auricles, auricles 1–2 mm;

petals light violet on both surfaces, lower 3 purple-veined, all beardless, lowest petal 8–20 mm, spur white, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers on prostrate to ascending peduncles.

Capsules

ellipsoid-ovoid, 6–16 mm, glabrous.

oblong, 8–10 mm, glabrous.

Seeds

light brown, blotched or streaked with brown, shiny, 2.1–2.7 mm.

dark brown to black, 1.2–2 mm.

2n

= 12.

= 44.

Viola lobata

Viola clauseniana

Phenology Flowering Apr–Jun.
Habitat Hanging gardens, seeps, springs, shady areas
Elevation 1600 m (5200 ft)
Distribution
from FNA
CA; OR; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

V. B. Baird (1942) noted that Viola lobata was more closely related to V. tripartita than to any western Viola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Viola clauseniana is endemic to Zion National Park, Washington County. M. S. Baker reported that its seeds were minutely roughed, a characteristic not recorded for other Viola species. Viola clauseniana was originally thought to be closely related to the acaulescent blue violets, most notably V. nephrophylla (S. L. Welsh et al. 1987; L. E. McKinney 1992). After contemplating the 2n = 44 chromosome count obtained by J. Clausen (1964), H. E. Ballard (pers. comm.) suggested that V. clauseniana might be more closely related with the stemless white violets (for example, V. blanda) than with V. nephrophylla. Viola clauseniana was considered a distinct species by N. H. Holmgren (2005d) and T. Marcussen and T. Karlsson (2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cauline leaf blades usually deeply divided, seldom unlobed, reniform, ± ovate, or deltate, if divided, palmately 3–12-lobed, lobe margins usually entire.
var. lobata
1. Cauline leaf blades unlobed, deltate to rhombic, ovate, or reniform-cordate, margins ± serrate, crenate-serrate, or dentate to deeply serrate or coarsely lacerate, often entire distally.
var. integrifolia
Source FNA vol. 6, p. 137. FNA vol. 6, p. 128.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. lobata var. integrifolia, V. lobata var. lobata
Name authority Bentham: Pl. Hartw., 298. (1849) M. S. Baker: Madroño 4: 194. (1938)
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