Viola lobata |
Viola brittoniana |
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moose horn violet, pine violet, yellow wood violet |
coast violet, coastal violet, northern coastal violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–46 cm. | Plants perennial, acaulescent, not stoloniferous, 5–30 cm; rhizome thick, fleshy. | ||||
Stems | 1–3, erect, leafless proximally, leafy distally, glabrous or puberulent, from subligneous rhizome. |
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Leaves | basal and cauline; basal: 0–2, unlobed or palmately 3–11-lobed; stipules ovate to lanceolate, margins ± entire or serrate, apex acute; petiole 5–27 cm, glabrous or puberulent; blade sometimes glaucous, deltate to reniform, 3.5–8.5 × 4.5–13.5 cm, base cordate, truncate, or attenuate, margins on unlobed leaves coarsely dentate-serrate, margins on lobed leaves usually entire, sometimes few-toothed, ciliate (sometimes only proximal 1/2) or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: distal on naked stems, unlobed and divided leaves can occur on same plant, if divided, palmately 3–12-lobed; stipules sometimes large and ± leaflike, margins entire, lacerate, or laciniate, sometimes with gland-tipped projections, apex acute to acuminate; petiole 0.2–8.8 cm; blade reniform to reniform-cordate, ± ovate, deltate, or rhombic, 1.5–5.5 × 1.4–10 cm, base cordate, subcordate, truncate, or attenuate, margins entire, crenate-serrate, or dentate, or coarsely lacerate to deeply serrate, often entire distally, ciliate or rarely eciliate, apex acute to obtuse, often long-tapered, mucronulate. |
basal, 5–9, ascending to erect, 5–9-lobed; stipules linear-lanceolate, margins entire, apex obtuse; petiole 3–16 cm, usually glabrous; mid-season blades incised or lobed throughout, earliest blades lobed (plants homophyllous), similar to mid-season blades, middle and lateral blade lobes differ in width and/or shape, middle lobes lanceolate or spatulate to narrowly obovate, lateral lobes lanceolate or spatulate to falcate (each sometimes with deltate to falcate appendages or teeth along margins), 1–7 × 2–8 cm, base truncate to cordate, margins entire, ciliate, apex acute to obtuse, surfaces usually glabrous, rarely with a few strigose, hairs concentrated on veins. |
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Peduncles | 2–13 cm, glabrous or pubescent. |
5–18 cm, usually glabrous. |
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Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 0.1–1 mm; petals deep lemon-yellow adaxially, usually upper 2 and sometimes lateral 2 brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined basally, lateral 2 bearded, lowest 8–19 mm, spur yellow to greenish, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 2–3 mm; petals light to soft reddish violet on both surfaces, lower 3 white basally, dark violet-veined, lateral 2 bearded, lowest 10–25 mm, sometimes bearded, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on ascending to erect peduncles. |
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Capsules | ellipsoid-ovoid, 6–16 mm, glabrous. |
ellipsoid, 10–15 mm, glabrous. |
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Seeds | light brown, blotched or streaked with brown, shiny, 2.1–2.7 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
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2n | = 12. |
= 54. |
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Viola lobata |
Viola brittoniana |
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Phenology | Flowering Apr–Jun. | |||||
Habitat | Alluvial woods, mesic forests on slopes near streams, wet fields, salt meadows | |||||
Elevation | 0–100 m (0–300 ft) | |||||
Distribution |
CA; OR; nw Mexico
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CT; DE; MA; MD; ME; NC; NJ; NY; PA; SC; VA |
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Discussion | Varieties 2 (2 in the flora). V. B. Baird (1942) noted that Viola lobata was more closely related to V. tripartita than to any western Viola. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Britton considered Viola brittoniana similar to V. pedatifida, apparently making him the first to recognize their affinity. Both are homophyllous and similar in other characters (L. E. McKinney 1992). McKinney recognized V. brittoniana as a variety of V. pedatifida. Additional evidence (N. L. Gil-Ad 1997; A. Haines 2011) suggests that these taxa are sufficiently distinct to continue recognition as separate species. Viola pectinata has sharply dentate leaves and is closely related to V. brittoniana, usually occurring with it. N. L. Gil-Ad (1997) made a convincing argument to recognize it as a form; A. Haines (2011) recognized it as a species. Others have considered it to be a sporadic form that may be of hybrid origin, or, as N. H. Russell (1965) suggested, is due to genetic dimorphism. We are reserving a decision on treating this taxon until additional studies are completed. A. Haines (2011) reported that the distribution of Viola brittoniana in Maine is actually based on the hybrid V. ×insolita House (V. brittoniana × V. sororia). Viola brittoniana reportedly hybridizes with V. cucullata (= V. ×notabilis E. P. Bicknell), V. sagittata var. sagittata (= V. ×mulfordiae Pollard), and V. sororia (= V. ×insolita). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 137. | FNA vol. 6, p. 125. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. atlantica, V. baxteri, V. brittoniana var. pectinata, V. pectinata, V. pedatifida subsp. brittoniana, V. pedatifida var. brittoniana | |||||
Name authority | Bentham: Pl. Hartw., 298. (1849) | Pollard: Bot. Gaz. 26: 332. (1898) | ||||
Web links |