Viola lobata |
Viola beckwithii |
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moose horn violet, pine violet, yellow wood violet |
Beckwith's or Great Basin violet, Beckwith's violet, Great Basin violet, sagebrush pansy, sagebrush violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–46 cm. | Plants perennial, caulescent, not stoloniferous, 2–22 cm. | ||||
Stems | 1–3, erect, leafless proximally, leafy distally, glabrous or puberulent, from subligneous rhizome. |
1–3, decumbent, ascending, or erect, ca. 1/2 subterranean, glabrous or usually puberulent, on single, short, vertical, deep-seated caudex. |
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Leaves | basal and cauline; basal: 0–2, unlobed or palmately 3–11-lobed; stipules ovate to lanceolate, margins ± entire or serrate, apex acute; petiole 5–27 cm, glabrous or puberulent; blade sometimes glaucous, deltate to reniform, 3.5–8.5 × 4.5–13.5 cm, base cordate, truncate, or attenuate, margins on unlobed leaves coarsely dentate-serrate, margins on lobed leaves usually entire, sometimes few-toothed, ciliate (sometimes only proximal 1/2) or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: distal on naked stems, unlobed and divided leaves can occur on same plant, if divided, palmately 3–12-lobed; stipules sometimes large and ± leaflike, margins entire, lacerate, or laciniate, sometimes with gland-tipped projections, apex acute to acuminate; petiole 0.2–8.8 cm; blade reniform to reniform-cordate, ± ovate, deltate, or rhombic, 1.5–5.5 × 1.4–10 cm, base cordate, subcordate, truncate, or attenuate, margins entire, crenate-serrate, or dentate, or coarsely lacerate to deeply serrate, often entire distally, ciliate or rarely eciliate, apex acute to obtuse, often long-tapered, mucronulate. |
basal and cauline; basal: 1–6 per caudex, palmately compound, ± 2-ternate or 3-ternate, leaflets 3; stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute; petiole 2–10.5 cm, usually puberulent; blade ovate to deltate, 2.4–5 × 3.5–4.5 cm, base tapered, ultimate leaflets dissected into oblong, elliptic, lanceolate, or oblanceolate lobes 1–7 mm wide, lobe margins entire, usually ciliate, apex acute to obtuse, mucronulate, surfaces usually puberulent, seldom glabrous, abaxial surface without prominent vein parallel to each margin; cauline similar to basal except: stipules linear, apex acuminate; petiole 2–5.7 cm, usually puberulent, rarely glabrous; blade 1–2.7 × 1.5–3 cm. |
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Peduncles | 2–13 cm, glabrous or pubescent. |
1.5–10.6(–15.7) cm, usually puberulent, seldom glabrous. |
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Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 0.1–1 mm; petals deep lemon-yellow adaxially, usually upper 2 and sometimes lateral 2 brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined basally, lateral 2 bearded, lowest 8–19 mm, spur yellow to greenish, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins eciliate, auricles 0.1–1 mm; petals dark reddish violet on both surfaces, lower 3 usually lilac, rarely white or whitish, lateral 2 bearded, with yellow patch basally, dark reddish violet-veined, lowest 10–22 mm, with yellow patch, dark reddish violet-veined, spur whitish or yellowish, tinged purple, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers absent. |
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Capsules | ellipsoid-ovoid, 6–16 mm, glabrous. |
oblong-ovoid, 7–12 mm, glabrous. |
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Seeds | light brown, blotched or streaked with brown, shiny, 2.1–2.7 mm. |
brown, 3–4 mm. |
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2n | = 12. |
= 24. |
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Viola lobata |
Viola beckwithii |
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Phenology | Flowering Mar–May. | |||||
Habitat | Dry or moist places, among shrubs or beneath pines | |||||
Elevation | 900–2700 m (3000–8900 ft) | |||||
Distribution |
CA; OR; nw Mexico
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CA; ID; NV; OR; UT
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Discussion | Varieties 2 (2 in the flora). V. B. Baird (1942) noted that Viola lobata was more closely related to V. tripartita than to any western Viola. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In some populations of Viola beckwithii, the three lower petals are white with a yellow area proximally (V. B. Baird 1942). Leaves have been described as palmately biternate or triternate (L. Abrams and R. S. Ferris 1923–1960, vol. 3), ternately decompound into linear segments (C. L. Hitchcock et al. 1955–1969, vol. 3), palmately three-parted then bipinnately parted into ultimate linear or spatulate segments (P. A. Munz 1959), and palmately about three times three-parted into linear or spatulate-linear segments (W. L. Jepson 1951). Some populations in northern California are nearly or completely glabrous, which M. S. Baker recognized as var. glabrata. Viola beckwithii is reported to hybridize with V. utahensis (G. Davidse 1976). Observed pollinators of V. beckwithii in Utah include Apis mellifera Linnaeus and Anthophora ursina Cresson (Davidse). When Cottam described Viola bonnevillensis, he suggested that it could be a hybrid between V. beckwithii and V. utahensis, and G. Davidse (1976) concurred. The type specimen of V. bonnevillensis (Cottam 7067, UT) was examined by R. J. Little. Because no similar forms are known to have been collected since 1939, it is presumed that this taxon is a hybrid. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 137. | FNA vol. 6, p. 122. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. beckwithii var. cachensis, V. beckwithii subsp. glabrata, V. bonnevillensis | |||||
Name authority | Bentham: Pl. Hartw., 298. (1849) | Torrey: in E. M. Durand, Pl. Pratten. Calif., 82. (1855) | ||||
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