Viola lobata |
Viola adunca |
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moose horn violet, pine violet, yellow wood violet |
Cascades early blue violet, common periwinkle, early blue violet, hook-spur violet, hookedspur violet, large periwinkle, sand violet, western dog violet, wild dog violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–46 cm. | Plants perennial, caulescent, not stoloniferous, 1.8–30(–35) cm. | ||||||||
Stems | 1–3, erect, leafless proximally, leafy distally, glabrous or puberulent, from subligneous rhizome. |
1–5, erect, ascending, or decumbent, sometimes later reclining to nearly prostrate, glabrous or puberulent, on caudex from subligneous rhizome. |
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Leaves | basal and cauline; basal: 0–2, unlobed or palmately 3–11-lobed; stipules ovate to lanceolate, margins ± entire or serrate, apex acute; petiole 5–27 cm, glabrous or puberulent; blade sometimes glaucous, deltate to reniform, 3.5–8.5 × 4.5–13.5 cm, base cordate, truncate, or attenuate, margins on unlobed leaves coarsely dentate-serrate, margins on lobed leaves usually entire, sometimes few-toothed, ciliate (sometimes only proximal 1/2) or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: distal on naked stems, unlobed and divided leaves can occur on same plant, if divided, palmately 3–12-lobed; stipules sometimes large and ± leaflike, margins entire, lacerate, or laciniate, sometimes with gland-tipped projections, apex acute to acuminate; petiole 0.2–8.8 cm; blade reniform to reniform-cordate, ± ovate, deltate, or rhombic, 1.5–5.5 × 1.4–10 cm, base cordate, subcordate, truncate, or attenuate, margins entire, crenate-serrate, or dentate, or coarsely lacerate to deeply serrate, often entire distally, ciliate or rarely eciliate, apex acute to obtuse, often long-tapered, mucronulate. |
basal and cauline; basal: 1–4; stipules linear to linear-lanceolate, margins entire or laciniate with gland-tipped projections, apex acute to acuminate; petiole 0.5–13.5 cm, glabrous or puberulent; blade usually ovate or ovate-deltate to ovate-orbiculate, sometimes ± reniform or oblong, 0.5–6.9 × 0.4–5 cm, base cordate, subcordate, truncate, or attenuate, usually decurrent on petiole, margins crenate to crenulate or entire, ciliate or eciliate, apex acute to obtuse, surfaces glabrous or sparsely to densely puberulent; cauline similar to basal except: stipule margins lacerate to laciniate; petiole 0.5–6.5 cm; blade 0.6–5.5 × 0.4–4.7 cm. |
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Peduncles | 2–13 cm, glabrous or pubescent. |
1.7–13.8 cm, glabrous or puberulent. |
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Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 0.1–1 mm; petals deep lemon-yellow adaxially, usually upper 2 and sometimes lateral 2 brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined basally, lateral 2 bearded, lowest 8–19 mm, spur yellow to greenish, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins ciliate or eciliate, auricles not enlarged in fruit, 0.5–2 mm; petals light- to deep- to lavender-violet on both surfaces, rarely white, lower 3 usually white basally, dark violet-veined, lateral 2 (and sometimes upper 2) bearded, lowest 7–17(–23) mm, spur purple to violet or white, elongated, 5–7 mm, tip straight or pointed, curved up or lateral; style head sparsely to densely bearded, sometimes beardless; cleistogamous flowers axillary. |
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Capsules | ellipsoid-ovoid, 6–16 mm, glabrous. |
short-ovoid, 6–11 mm, glabrous. |
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Seeds | light brown, blotched or streaked with brown, shiny, 2.1–2.7 mm. |
dark brown to olive-black, 1.5–2 mm. |
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2n | = 12. |
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Viola lobata |
Viola adunca |
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Distribution |
CA; OR; nw Mexico
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AK; AZ; CA; CO; IA; ID; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OR; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NS; NT; NU; ON; QC; SK; YT
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Discussion | Varieties 2 (2 in the flora). V. B. Baird (1942) noted that Viola lobata was more closely related to V. tripartita than to any western Viola. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Viola adunca is polymorphic with over 50 named taxa. G. D. McPherson and J. G. Packer (1974) reported that diploid, triploid, and tetraploid races of V. adunca occur in Canada and northwestern United States. They found that diploid and tetraploid chromosome races can be distinguished morphologically based on style beards and on the size of guard cells and pollen grains and recommended taxonomic recognition of both races. In diploid races, the style projections are more or less cylindrical and about one-sixth the width of the style; in tetraploid races they are short-conical or globular, and about one-tenth or less than the width of the style beard. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 137. | FNA vol. 6, p. 119. | ||||||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Lophion aduncum, V. canina var. adunca | |||||||||
Name authority | Bentham: Pl. Hartw., 298. (1849) | Smith: in A. Rees, Cycl. 37: Viola no. 63 | ||||||||
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