Viola langsdorffii |
Viola sagittata |
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Alaska violet, Aleutian violet, Langsdorff's violet, violette de Langsdorff |
arrow-leaf violet, arrowhead violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 2–30 cm. | Plants perennial, acaulescent, not stoloniferous, 5–50 cm; rhizome thick, fleshy. | ||||
Stems | 1–3, ascending to erect, usually glabrous, from shallow, fleshy rhizome. |
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Leaves | basal and cauline; basal: 2–3; stipules broadly ovate to deltate or long-lanceolate, margins often glandular (glands sessile or stalked), apex acute; petiole 0.8–21 cm, glabrous or sparsely pubescent; blade ovate to reniform, 0.9–5.8 × 1–6 cm, base cordate to ± truncate, margins crenate to crenate-serrulate, usually eciliate, apex rounded or acute to usually obtuse, surfaces glabrous or sparsely pubescent on adaxial veins, seldom on abaxial veins; cauline similar to basal except: stipules broadly ovate to lanceolate-oblong or deltate, apex acute to acuminate; petiole 2.2–12.1 cm; blade 1.9–4.2 × 1.8–5.9 cm. |
basal, 4–8, ascending to erect; stipules linear-lanceolate, margins entire, apex acute; petiole 2–13 cm, glabrous or pubescent; middle and lateral blade lobes differ in width and/or shape, earliest blades ovate to elliptic, mid-season blades ovate, elliptic, narrowly elliptic to narrowly deltate, 1–8 × 1–5 cm, incised or lobed at base only, base sagittate or hastate, truncate, attenuate, or ± cordate, margins crenate or serrate, ciliate or eciliate, apex acute, surfaces glabrous or pubescent. |
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Peduncles | 2.2–20.7 cm, glabrous or sparsely pubescent. |
3–15 cm, glabrous or pubescent. |
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Flowers | sepals lanceolate to ovate, margins eciliate, auricles 0.5–1.5 mm; petals light to deep violet on both surfaces, lower 3 white basally and light to deep violet-veined, lateral 2 bearded, lowest 12–24 mm, spur white to violet, gibbous to elongated, 2–5 mm, usually less than 1/2 lowest petal, tip straight; style head usually beardless, sometimes bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 2–3 mm; petals light to dark violet on both surfaces, lower 3 white basally, lowest dark violet-veined, lateral 2 densely bearded, spur on lowest petal occasionally bearded, lowest 10–15 mm, spur light to dark violet, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate, ascending, or erect peduncles. |
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Capsules | ovoid to oblong, 7.5–13 mm, glabrous. |
ellipsoid, 10–14 mm, glabrous. |
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Seeds | dark olive to ± black, 2.5–2.8 mm. |
beige, mottled to bronze, 1.3–2.5 mm. |
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2n | = 12. |
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Viola langsdorffii |
Viola sagittata |
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Phenology | Flowering Apr–Aug. | |||||
Habitat | Swamps, bogs, fens, wet meadows, stream banks, rocky slopes, usually near coast | |||||
Elevation | 0–1500 m (0–4900 ft) | |||||
Distribution |
AK; CA; OR; WA; BC; YT; Asia
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AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; NB; NS; ON; PE; QC
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Discussion | Viola simulata and V. superba are closely related to V. langsdorffii (L. Abrams and R. S. Ferris 1923–1960, vol. 3). They were treated by C. L. Hitchcock et al. (1955–1969, vol. 3) as synonymous with V. langsdorffii pending further study. E. Brainerd (1921) stated that Viola langsdorffii presents an interesting transition between the caulescent and acaulescent violets. Plants of the species are reported from Winchester Grade in Idaho (Gail s.n., 1931, ID); verification is needed. Viola langsdorffii is reported to have fleshy rhizomes. Plants in Del Norte County, California, have fleshy rhizomes near the soil surface, but these are connected to deeper, subligneous rhizomes. The style in some populations of Viola langsdorffii is reported to be sometimes bearded (G. W. Douglas et al. 1998–2002, vol. 5; W. J. Cody 2000). Illustrations in V. B. Baird (1942; location of plant unknown) and M. S. Baker (1936; a plant from Alaska) show a beardless style; L. Abrams and R. S. Ferris (1923–1960, vol. 3) and C. L. Hitchcock et al. (1955–1969, vol. 3) reported that the style head was beardless. Styles are beardless in plants occurring in Del Norte County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 136. | FNA vol. 6, p. 155. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. simulata, V. superba | |||||
Name authority | Fischer ex Gingins: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 1: 296. (1824) | Aiton: Hort. Kew. 3: 287. (1789) | ||||
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