Viola langsdorffii |
Viola rotundifolia |
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Alaska violet, Aleutian violet, Langsdorff's violet, violette de Langsdorff |
early yellow or roundleaf yellow violet, round-leaf violet, roundleaf yellow violet, violette à feuilles rondes |
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Habit | Plants perennial, caulescent, not stoloniferous, 2–30 cm. | Plants perennial, acaulescent, not stoloniferous, 1–20 cm; rhizome thick, fleshy. |
Stems | 1–3, ascending to erect, usually glabrous, from shallow, fleshy rhizome. |
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Leaves | basal and cauline; basal: 2–3; stipules broadly ovate to deltate or long-lanceolate, margins often glandular (glands sessile or stalked), apex acute; petiole 0.8–21 cm, glabrous or sparsely pubescent; blade ovate to reniform, 0.9–5.8 × 1–6 cm, base cordate to ± truncate, margins crenate to crenate-serrulate, usually eciliate, apex rounded or acute to usually obtuse, surfaces glabrous or sparsely pubescent on adaxial veins, seldom on abaxial veins; cauline similar to basal except: stipules broadly ovate to lanceolate-oblong or deltate, apex acute to acuminate; petiole 2.2–12.1 cm; blade 1.9–4.2 × 1.8–5.9 cm. |
basal, 2–5, prostrate to ascending, often overlapping basally; stipules linear-lanceolate, margins entire, apex acute; petiole 2–8 cm, pubescent; blade unlobed, orbiculate, reniform, or ovate, 2–12 × 1.5–9 cm, base cordate, margins crenate to serrate, sometimes glandular, ciliate or eciliate, apex rounded to acute, surfaces usually pubescent throughout or concentrated proximally on both surfaces. |
Peduncles | 2.2–20.7 cm, glabrous or sparsely pubescent. |
1.5–7 cm, usually pubescent. |
Flowers | sepals lanceolate to ovate, margins eciliate, auricles 0.5–1.5 mm; petals light to deep violet on both surfaces, lower 3 white basally and light to deep violet-veined, lateral 2 bearded, lowest 12–24 mm, spur white to violet, gibbous to elongated, 2–5 mm, usually less than 1/2 lowest petal, tip straight; style head usually beardless, sometimes bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 0.5–1 mm; petals deep lemon-yellow on both surfaces, lower 3 brownish purple-veined, lateral 2 bearded, lowest 8–11 mm, spur yellow, gibbous, 1–2 mm; style head beardless; cleistogamous flowers on prostrate or partially subterranean rhizomes or on racemelike, nonrooting, and usually leafless branches growing from rhizome apex. |
Capsules | ovoid to oblong, 7.5–13 mm, glabrous. |
ellipsoid, 5–10 mm, glabrous. |
Seeds | dark olive to ± black, 2.5–2.8 mm. |
beige, 1–2 mm. |
2n | = 12. |
= 12. |
Viola langsdorffii |
Viola rotundifolia |
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Phenology | Flowering Apr–Aug. | Flowering Mar–May. |
Habitat | Swamps, bogs, fens, wet meadows, stream banks, rocky slopes, usually near coast | Rich montane forests and other mesic woodlands |
Elevation | 0–1500 m (0–4900 ft) | 200–2000 m (700–6600 ft) |
Distribution |
AK; CA; OR; WA; BC; YT; Asia
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CT; DE; GA; KY; MA; MD; ME; NC; NH; NY; OH; PA; RI; SC; TN; VA; VT; WV; ON; QC
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Discussion | Viola simulata and V. superba are closely related to V. langsdorffii (L. Abrams and R. S. Ferris 1923–1960, vol. 3). They were treated by C. L. Hitchcock et al. (1955–1969, vol. 3) as synonymous with V. langsdorffii pending further study. E. Brainerd (1921) stated that Viola langsdorffii presents an interesting transition between the caulescent and acaulescent violets. Plants of the species are reported from Winchester Grade in Idaho (Gail s.n., 1931, ID); verification is needed. Viola langsdorffii is reported to have fleshy rhizomes. Plants in Del Norte County, California, have fleshy rhizomes near the soil surface, but these are connected to deeper, subligneous rhizomes. The style in some populations of Viola langsdorffii is reported to be sometimes bearded (G. W. Douglas et al. 1998–2002, vol. 5; W. J. Cody 2000). Illustrations in V. B. Baird (1942; location of plant unknown) and M. S. Baker (1936; a plant from Alaska) show a beardless style; L. Abrams and R. S. Ferris (1923–1960, vol. 3) and C. L. Hitchcock et al. (1955–1969, vol. 3) reported that the style head was beardless. Styles are beardless in plants occurring in Del Norte County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
N. H. Russell (1955b) stated that Viola rotundifolia is a primitive member of Viola and probably one of the ancestral species of stemmed yellow violets of North America. Russell (1965) stated that morphologically, V. rotundifolia is one of the most invariable violets and suggested that its nearest relative is V. orbiculata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 136. | FNA vol. 6, p. 155. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. simulata, V. superba | |
Name authority | Fischer ex Gingins: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 1: 296. (1824) | Michaux: Fl. Bor.-Amer. 2: 150. (1803) |
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