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Alaska violet, Aleutian violet, Langsdorff's violet, violette de Langsdorff

primrose-leaf violet, western bog violet, western white bog violet

Habit Plants perennial, caulescent, not stoloniferous, 2–30 cm. Plants perennial, acaulescent, stoloniferous, (3–)5–20(–36) cm; stolons pale, often rooting and leafy at nodes; rhizome thick or slender, fleshy.
Stems

1–3, ascending to erect, usually glabrous, from shallow, fleshy rhizome.

Leaves

basal and cauline;

basal: 2–3;

stipules broadly ovate to deltate or long-lanceolate, margins often glandular (glands sessile or stalked), apex acute;

petiole 0.8–21 cm, glabrous or sparsely pubescent;

blade ovate to reniform, 0.9–5.8 × 1–6 cm, base cordate to ± truncate, margins crenate to crenate-serrulate, usually eciliate, apex rounded or acute to usually obtuse, surfaces glabrous or sparsely pubescent on adaxial veins, seldom on abaxial veins;

cauline similar to basal except: stipules broadly ovate to lanceolate-oblong or deltate, apex acute to acuminate;

petiole 2.2–12.1 cm;

blade 1.9–4.2 × 1.8–5.9 cm.

basal, 4–8, prostrate to ascending;

stipules linear-lanceolate, margins ± crenate-serrate, sometimes glandular, apex acute;

petiole (1–)3–13(–29) cm, glabrous or pubescent;

blade unlobed, elliptic to narrowly or broadly ovate, (1.5–)3–7(–9) × 1–3(–3.5) cm, longer than wide, base broadly cordate to attenuate, rarely ± truncate, margins crenulate to serrulate, sometimes glandular, mostly eciliate, apex acute to rounded, mucronulate, surfaces glabrous or sparsely pubescent.

Peduncles

2.2–20.7 cm, glabrous or sparsely pubescent.

(3–)6–18(–28) cm, glabrous.

Flowers

sepals lanceolate to ovate, margins eciliate, auricles 0.5–1.5 mm;

petals light to deep violet on both surfaces, lower 3 white basally and light to deep violet-veined, lateral 2 bearded, lowest 12–24 mm, spur white to violet, gibbous to elongated, 2–5 mm, usually less than 1/2 lowest petal, tip straight;

style head usually beardless, sometimes bearded; cleistogamous flowers axillary.

sepals lanceolate to ovate, margins usually eciliate, auricles 1–2 mm;

petals white on both surfaces, lower 3 purple-veined, lateral 2 bearded or beardless, lowest 9–14(–16) mm, spur white, gibbous, 1–2 mm;

style head beardless; cleistogamous flowers present or absent.

Capsules

ovoid to oblong, 7.5–13 mm, glabrous.

ellipsoid, (5–)8–9 mm, glabrous.

Seeds

dark olive to ± black, 2.5–2.8 mm.

beige to bronze, 1.5–2 mm.

2n

= 12.

= 24.

Viola langsdorffii

Viola primulifolia

Phenology Flowering Apr–Aug.
Habitat Swamps, bogs, fens, wet meadows, stream banks, rocky slopes, usually near coast
Elevation 0–1500 m (0–4900 ft)
Distribution
from FNA
AK; CA; OR; WA; BC; YT; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; CT; DC; DE; FL; GA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MS; NC; NH; NJ; NY; OH; OK; OR; PA; RI; SC; TN; TX; VA; VT; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola simulata and V. superba are closely related to V. langsdorffii (L. Abrams and R. S. Ferris 1923–1960, vol. 3). They were treated by C. L. Hitchcock et al. (1955–1969, vol. 3) as synonymous with V. langsdorffii pending further study.

E. Brainerd (1921) stated that Viola langsdorffii presents an interesting transition between the caulescent and acaulescent violets. Plants of the species are reported from Winchester Grade in Idaho (Gail s.n., 1931, ID); verification is needed. Viola langsdorffii is reported to have fleshy rhizomes. Plants in Del Norte County, California, have fleshy rhizomes near the soil surface, but these are connected to deeper, subligneous rhizomes.

The style in some populations of Viola langsdorffii is reported to be sometimes bearded (G. W. Douglas et al. 1998–2002, vol. 5; W. J. Cody 2000). Illustrations in V. B. Baird (1942; location of plant unknown) and M. S. Baker (1936; a plant from Alaska) show a beardless style; L. Abrams and R. S. Ferris (1923–1960, vol. 3) and C. L. Hitchcock et al. (1955–1969, vol. 3) reported that the style head was beardless. Styles are beardless in plants occurring in Del Norte County, California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Viola primulifolia occurs in small colonies; individual plants are interconnected by stolons.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades elliptic to widely ovate, base broadly cordate to attenuate, surfaces sparsely pubescent, rarely glabrous; cleistogamous flowers present; c, e United States.
var. primulifolia
1. Leaf blades elliptic to narrowly ovate, base usually attenuate, rarely ± truncate, surfaces glabrous, rarely pubescent; cleistogamous flowers absent; nw California, sw Oregon.
var. occidentalis
Source FNA vol. 6, p. 136. FNA vol. 6, p. 148.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. primulifolia var. occidentalis, V. primulifolia var. primulifolia
Synonyms V. simulata, V. superba
Name authority Fischer ex Gingins: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 1: 296. (1824) Linnaeus: Sp. Pl. 2: 934. (1753)
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