Viola langsdorffii |
Viola pinetorum |
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Alaska violet, Aleutian violet, Langsdorff's violet, violette de Langsdorff |
goosefoot yellow violet, mountain yellow violet, pine violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 2–30 cm. | Plants perennial, caulescent, not stoloniferous, 3–18 cm, cespitose or not. | ||||
Stems | 1–3, ascending to erect, usually glabrous, from shallow, fleshy rhizome. |
1–3, prostrate or erect, leafy proximally and distally, puberulent or canescent to gray-tomentose, sometimes glabrous, on caudex from subligneous rhizome. |
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Leaves | basal and cauline; basal: 2–3; stipules broadly ovate to deltate or long-lanceolate, margins often glandular (glands sessile or stalked), apex acute; petiole 0.8–21 cm, glabrous or sparsely pubescent; blade ovate to reniform, 0.9–5.8 × 1–6 cm, base cordate to ± truncate, margins crenate to crenate-serrulate, usually eciliate, apex rounded or acute to usually obtuse, surfaces glabrous or sparsely pubescent on adaxial veins, seldom on abaxial veins; cauline similar to basal except: stipules broadly ovate to lanceolate-oblong or deltate, apex acute to acuminate; petiole 2.2–12.1 cm; blade 1.9–4.2 × 1.8–5.9 cm. |
basal and cauline; basal: 1–4; stipules adnate to petiole forming 2 linear-lanceolate wings, margins entire or laciniate, apex of each wing free, tips usually filamentous; petiole 2.3–9.5 cm, puberulent or canescent; blade purple-tinted abaxially or not, usually linear to narrowly lanceolate, oblanceolate or obovate, or lanceolate-elliptic, rarely ovate, 1.3–5 × 0.3–2.5 cm, base attenuate, margins usually lacerate, dentate, or serrate, sometimes entire, usually undulate, ciliate, apex acute, mucronulate, surfaces puberulent to canescent or gray-tomentose; cauline similar to basal except: stipules lanceolate, oblanceolate, or linear-oblong, margins entire or lacerate, apex acute to acuminate; petiole 0.9–8.3 cm; blade 2.8–9.6 × 0.3–1.4 cm, length 4–11 times width. |
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Peduncles | 2.2–20.7 cm, glabrous or sparsely pubescent. |
2.9–11.5 cm, puberulent or canescent. |
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Flowers | sepals lanceolate to ovate, margins eciliate, auricles 0.5–1.5 mm; petals light to deep violet on both surfaces, lower 3 white basally and light to deep violet-veined, lateral 2 bearded, lowest 12–24 mm, spur white to violet, gibbous to elongated, 2–5 mm, usually less than 1/2 lowest petal, tip straight; style head usually beardless, sometimes bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins ciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 red- to purple-brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 5–12 mm, spur color same as petals, gibbous, 1.5–3 mm; style head bearded; cleistogamous flowers axillary. |
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Capsules | ovoid to oblong, 7.5–13 mm, glabrous. |
ovoid, 3.5–7 mm, puberulent. |
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Seeds | dark olive to ± black, 2.5–2.8 mm. |
medium to dark brown, 2–3.5 mm. |
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2n | = 12. |
= 12. |
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Viola langsdorffii |
Viola pinetorum |
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Phenology | Flowering Apr–Aug. | |||||
Habitat | Swamps, bogs, fens, wet meadows, stream banks, rocky slopes, usually near coast | |||||
Elevation | 0–1500 m (0–4900 ft) | |||||
Distribution |
AK; CA; OR; WA; BC; YT; Asia
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CA
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Discussion | Viola simulata and V. superba are closely related to V. langsdorffii (L. Abrams and R. S. Ferris 1923–1960, vol. 3). They were treated by C. L. Hitchcock et al. (1955–1969, vol. 3) as synonymous with V. langsdorffii pending further study. E. Brainerd (1921) stated that Viola langsdorffii presents an interesting transition between the caulescent and acaulescent violets. Plants of the species are reported from Winchester Grade in Idaho (Gail s.n., 1931, ID); verification is needed. Viola langsdorffii is reported to have fleshy rhizomes. Plants in Del Norte County, California, have fleshy rhizomes near the soil surface, but these are connected to deeper, subligneous rhizomes. The style in some populations of Viola langsdorffii is reported to be sometimes bearded (G. W. Douglas et al. 1998–2002, vol. 5; W. J. Cody 2000). Illustrations in V. B. Baird (1942; location of plant unknown) and M. S. Baker (1936; a plant from Alaska) show a beardless style; L. Abrams and R. S. Ferris (1923–1960, vol. 3) and C. L. Hitchcock et al. (1955–1969, vol. 3) reported that the style head was beardless. Styles are beardless in plants occurring in Del Norte County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Flowers of Viola pinetorum have been observed to close up in late afternoon then fully reopen the following morning. Although E. O. Wooton and P. C. Standley (1915) reported Viola pinetorum from New Mexico, the plant was probably V. nuttallii. K. W. Allred (2008) noted that V. pinetorum occurs in California; he did not recognize it in New Mexico. Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 136. | FNA vol. 6, p. 144. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. simulata, V. superba | V. purpurea var. pinetorum | ||||
Name authority | Fischer ex Gingins: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 1: 296. (1824) | Greene: Pittonia 2: 14. (1889) | ||||
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