Viola langsdorffii |
Viola pedata |
|||||
---|---|---|---|---|---|---|
Alaska violet, Aleutian violet, Langsdorff's violet, violette de Langsdorff |
bird-foot violet |
|||||
Habit | Plants perennial, caulescent, not stoloniferous, 2–30 cm. | Plants perennial, acaulescent, not stoloniferous, 5–30 cm; rhizome thick, fleshy. | ||||
Stems | 1–3, ascending to erect, usually glabrous, from shallow, fleshy rhizome. |
|||||
Leaves | basal and cauline; basal: 2–3; stipules broadly ovate to deltate or long-lanceolate, margins often glandular (glands sessile or stalked), apex acute; petiole 0.8–21 cm, glabrous or sparsely pubescent; blade ovate to reniform, 0.9–5.8 × 1–6 cm, base cordate to ± truncate, margins crenate to crenate-serrulate, usually eciliate, apex rounded or acute to usually obtuse, surfaces glabrous or sparsely pubescent on adaxial veins, seldom on abaxial veins; cauline similar to basal except: stipules broadly ovate to lanceolate-oblong or deltate, apex acute to acuminate; petiole 2.2–12.1 cm; blade 1.9–4.2 × 1.8–5.9 cm. |
basal, 4–10, ascending to erect, deeply divided; stipules linear-lanceolate, margins entire, lacerate, or shallowly divided, apex acute; petiole 2–12 cm, glabrous or sparsely puberulent; blade 3–9(–10)-lobed, lobes similar in width and shape, spatulate, lanceolate ± linear, deltate, or ovate, 1–4 × 1–4 cm, base attenuate or broadly cordate to cuneate, margins entire, ciliate or eciliate, apex rounded to usually acute, surfaces usually glabrous, sometimes pubescent on abaxial veins. |
||||
Peduncles | 2.2–20.7 cm, glabrous or sparsely pubescent. |
5–12 cm, glabrous or pubescent. |
||||
Flowers | sepals lanceolate to ovate, margins eciliate, auricles 0.5–1.5 mm; petals light to deep violet on both surfaces, lower 3 white basally and light to deep violet-veined, lateral 2 bearded, lowest 12–24 mm, spur white to violet, gibbous to elongated, 2–5 mm, usually less than 1/2 lowest petal, tip straight; style head usually beardless, sometimes bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins mostly ciliate, at least proximally, auricles 1–2 mm; petals uniformly light to dark blue-violet on both surfaces or upper 2 darker adaxially, sometimes white, upper and lateral 2 often darker basally, lowest, seldom others, dark violet-veined, all beardless, lowest white basally, 12–24 mm, spur white, gibbous, 2–3 mm; style head beardless; cleistogamous flowers absent. |
||||
Capsules | ovoid to oblong, 7.5–13 mm, glabrous. |
ellipsoid, 6–10 mm, glabrous. |
||||
Seeds | dark olive to ± black, 2.5–2.8 mm. |
beige, mottled to brown, 1.4–3 mm. |
||||
2n | = 12. |
= 56. |
||||
Viola langsdorffii |
Viola pedata |
|||||
Phenology | Flowering Apr–Aug. | |||||
Habitat | Swamps, bogs, fens, wet meadows, stream banks, rocky slopes, usually near coast | |||||
Elevation | 0–1500 m (0–4900 ft) | |||||
Distribution |
AK; CA; OR; WA; BC; YT; Asia
|
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; ON
|
||||
Discussion | Viola simulata and V. superba are closely related to V. langsdorffii (L. Abrams and R. S. Ferris 1923–1960, vol. 3). They were treated by C. L. Hitchcock et al. (1955–1969, vol. 3) as synonymous with V. langsdorffii pending further study. E. Brainerd (1921) stated that Viola langsdorffii presents an interesting transition between the caulescent and acaulescent violets. Plants of the species are reported from Winchester Grade in Idaho (Gail s.n., 1931, ID); verification is needed. Viola langsdorffii is reported to have fleshy rhizomes. Plants in Del Norte County, California, have fleshy rhizomes near the soil surface, but these are connected to deeper, subligneous rhizomes. The style in some populations of Viola langsdorffii is reported to be sometimes bearded (G. W. Douglas et al. 1998–2002, vol. 5; W. J. Cody 2000). Illustrations in V. B. Baird (1942; location of plant unknown) and M. S. Baker (1936; a plant from Alaska) show a beardless style; L. Abrams and R. S. Ferris (1923–1960, vol. 3) and C. L. Hitchcock et al. (1955–1969, vol. 3) reported that the style head was beardless. Styles are beardless in plants occurring in Del Norte County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||
Key |
|
|||||
Source | FNA vol. 6, p. 136. | FNA vol. 6, p. 143. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. simulata, V. superba | |||||
Name authority | Fischer ex Gingins: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 1: 296. (1824) | Linnaeus: Sp. Pl. 2: 933. (1753) | ||||
Web links |
|