Viola langsdorffii |
Viola egglestonii |
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Alaska violet, Aleutian violet, Langsdorff's violet, violette de Langsdorff |
cedar glade or glade violet, glade violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 2–30 cm. | Plants perennial, acaulescent, not stoloniferous, 5–20 cm; rhizome thick, fleshy. |
Stems | 1–3, ascending to erect, usually glabrous, from shallow, fleshy rhizome. |
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Leaves | basal and cauline; basal: 2–3; stipules broadly ovate to deltate or long-lanceolate, margins often glandular (glands sessile or stalked), apex acute; petiole 0.8–21 cm, glabrous or sparsely pubescent; blade ovate to reniform, 0.9–5.8 × 1–6 cm, base cordate to ± truncate, margins crenate to crenate-serrulate, usually eciliate, apex rounded or acute to usually obtuse, surfaces glabrous or sparsely pubescent on adaxial veins, seldom on abaxial veins; cauline similar to basal except: stipules broadly ovate to lanceolate-oblong or deltate, apex acute to acuminate; petiole 2.2–12.1 cm; blade 1.9–4.2 × 1.8–5.9 cm. |
basal, 3, prostrate to ascending; stipules linear-lanceolate, margins entire, apex acute; petiole 1.5–7 cm, usually glabrous; earliest leaf blades ± deltate or 3-lobed, mid-season blades 5–9-lobed, 1–9 × 1–10 cm, base truncate to cuneate, middle lobe lanceolate or spatulate to narrowly ovate, lateral lobes lanceolate or spatulate to falcate, margins serrate, sometimes with deltate or falcate appendages or teeth, ciliate, apex acute, surfaces usually glabrous, rarely pubescent. |
Peduncles | 2.2–20.7 cm, glabrous or sparsely pubescent. |
2–15 cm, usually glabrous, rarely pubescent. |
Flowers | sepals lanceolate to ovate, margins eciliate, auricles 0.5–1.5 mm; petals light to deep violet on both surfaces, lower 3 white basally and light to deep violet-veined, lateral 2 bearded, lowest 12–24 mm, spur white to violet, gibbous to elongated, 2–5 mm, usually less than 1/2 lowest petal, tip straight; style head usually beardless, sometimes bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 0.5–1 mm; petals light to dark blue-violet on both surfaces, lower 3 and sometimes upper 2 white basally, lower 3 darker violet-veined, lateral 2 densely bearded, lowest 10–15 mm, spur usually lilac, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate peduncles. |
Capsules | ovoid to oblong, 7.5–13 mm, glabrous. |
ellipsoid, 11–14 mm, glabrous. |
Seeds | dark olive to ± black, 2.5–2.8 mm. |
beige, mottled to bronze, 2–3 mm. |
2n | = 12. |
= 54. |
Viola langsdorffii |
Viola egglestonii |
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Phenology | Flowering Apr–Aug. | Flowering Mar–May. |
Habitat | Swamps, bogs, fens, wet meadows, stream banks, rocky slopes, usually near coast | Limestone glades and barrens |
Elevation | 0–1500 m (0–4900 ft) | 100–200 m (300–700 ft) |
Distribution |
AK; CA; OR; WA; BC; YT; Asia
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AL; GA; IN; KY; TN |
Discussion | Viola simulata and V. superba are closely related to V. langsdorffii (L. Abrams and R. S. Ferris 1923–1960, vol. 3). They were treated by C. L. Hitchcock et al. (1955–1969, vol. 3) as synonymous with V. langsdorffii pending further study. E. Brainerd (1921) stated that Viola langsdorffii presents an interesting transition between the caulescent and acaulescent violets. Plants of the species are reported from Winchester Grade in Idaho (Gail s.n., 1931, ID); verification is needed. Viola langsdorffii is reported to have fleshy rhizomes. Plants in Del Norte County, California, have fleshy rhizomes near the soil surface, but these are connected to deeper, subligneous rhizomes. The style in some populations of Viola langsdorffii is reported to be sometimes bearded (G. W. Douglas et al. 1998–2002, vol. 5; W. J. Cody 2000). Illustrations in V. B. Baird (1942; location of plant unknown) and M. S. Baker (1936; a plant from Alaska) show a beardless style; L. Abrams and R. S. Ferris (1923–1960, vol. 3) and C. L. Hitchcock et al. (1955–1969, vol. 3) reported that the style head was beardless. Styles are beardless in plants occurring in Del Norte County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
N. H. Russell (1965) was the first to consider Viola egglestonii similar to V. septemloba. After analyzing leaf-blade lobing patterns, L. E. McKinney considered it a subspecies of V. septemloba. Although similarities exist between these taxa, we maintain them as distinct species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 136. | FNA vol. 6, p. 129. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. simulata, V. superba | V. septemloba subsp. egglestonii |
Name authority | Fischer ex Gingins: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 1: 296. (1824) | Brainerd: Bull. Torrey Bot. Club 37: 526, plates 34, 35. (1910) |
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