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Alaska violet, Aleutian violet, Langsdorff's violet, violette de Langsdorff

Le Conte's violet, sand violet, violette affine

Habit Plants perennial, caulescent, not stoloniferous, 2–30 cm. Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age.
Stems

1–3, ascending to erect, usually glabrous, from shallow, fleshy rhizome.

Leaves

basal and cauline;

basal: 2–3;

stipules broadly ovate to deltate or long-lanceolate, margins often glandular (glands sessile or stalked), apex acute;

petiole 0.8–21 cm, glabrous or sparsely pubescent;

blade ovate to reniform, 0.9–5.8 × 1–6 cm, base cordate to ± truncate, margins crenate to crenate-serrulate, usually eciliate, apex rounded or acute to usually obtuse, surfaces glabrous or sparsely pubescent on adaxial veins, seldom on abaxial veins;

cauline similar to basal except: stipules broadly ovate to lanceolate-oblong or deltate, apex acute to acuminate;

petiole 2.2–12.1 cm;

blade 1.9–4.2 × 1.8–5.9 cm.

basal, 1–6, ascending to erect;

stipules lanceolate, margins entire or fimbriate, apex acute;

petiole 2–10 cm, glabrous;

blade green abaxially, unlobed, narrowly to broadly ovate or narrowly deltate, 1.5–10 × 1.5–10 cm, not fleshy, base cordate to broadly cordate or almost truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse, surfaces sparsely pubescent adaxially, rarely glabrous.

Peduncles

2.2–20.7 cm, glabrous or sparsely pubescent.

3–15 cm, glabrous or pubescent.

Flowers

sepals lanceolate to ovate, margins eciliate, auricles 0.5–1.5 mm;

petals light to deep violet on both surfaces, lower 3 white basally and light to deep violet-veined, lateral 2 bearded, lowest 12–24 mm, spur white to violet, gibbous to elongated, 2–5 mm, usually less than 1/2 lowest petal, tip straight;

style head usually beardless, sometimes bearded; cleistogamous flowers axillary.

sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm;

petals lavender-violet to dull reddish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, lowest 10–22 mm, usually obviously bearded, rarely beardless, spur white or same color as petals, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers from prostrate to ascending peduncles.

Capsules

ovoid to oblong, 7.5–13 mm, glabrous.

often reddish or purplish-flecked or green, ellipsoid, 5–10 mm, glabrous or puberulent.

Seeds

dark olive to ± black, 2.5–2.8 mm.

beige, mottled to bronze, 1.5–2.5 mm.

2n

= 12.

= 54.

Viola langsdorffii

Viola affinis

Phenology Flowering Apr–Aug. Flowering Apr–Jun.
Habitat Swamps, bogs, fens, wet meadows, stream banks, rocky slopes, usually near coast Open or wooded wet areas, meadows, stream banks, thickets, shores of lakes, seasonally dry areas.
Elevation 0–1500 m (0–4900 ft) 100–2000 m (300–6600 ft)
Distribution
from FNA
AK; CA; OR; WA; BC; YT; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; DC; DE; IL; IN; MA; MD; MI; MN; NH; NJ; NY; OH; PA; RI; VA; VT; WI; WV; ON; QC
[BONAP county map]
Discussion

Viola simulata and V. superba are closely related to V. langsdorffii (L. Abrams and R. S. Ferris 1923–1960, vol. 3). They were treated by C. L. Hitchcock et al. (1955–1969, vol. 3) as synonymous with V. langsdorffii pending further study.

E. Brainerd (1921) stated that Viola langsdorffii presents an interesting transition between the caulescent and acaulescent violets. Plants of the species are reported from Winchester Grade in Idaho (Gail s.n., 1931, ID); verification is needed. Viola langsdorffii is reported to have fleshy rhizomes. Plants in Del Norte County, California, have fleshy rhizomes near the soil surface, but these are connected to deeper, subligneous rhizomes.

The style in some populations of Viola langsdorffii is reported to be sometimes bearded (G. W. Douglas et al. 1998–2002, vol. 5; W. J. Cody 2000). Illustrations in V. B. Baird (1942; location of plant unknown) and M. S. Baker (1936; a plant from Alaska) show a beardless style; L. Abrams and R. S. Ferris (1923–1960, vol. 3) and C. L. Hitchcock et al. (1955–1969, vol. 3) reported that the style head was beardless. Styles are beardless in plants occurring in Del Norte County, California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

L. E. McKinney (1992) considered Viola affinis, and much of what botanists had called V. nephrophylla, to be essentially the same taxon. After studying additional specimens, reviewing literature (H. E. Ballard 1994; A. Haines 2001b), and discussions with others (J. Cayouette, H. E. Ballard, A. Haines, pers. comm.), he chose to maintain these as separate taxa. Reports of V. affinis in the Gulf coastal states based on specimens or photographs are usually attributable to V. missouriensis.

Viola affinis reportedly hybridizes with V. hirsutula (= V. ×consobrina House), V. cucullata (= V. ×consocia House), V. brittoniana (= V. ×davisii House), V. sororia (= V. ×filicetorum Greene [as species]), V. sagittata var. sagittata (= V. ×hollickii House), and V. nephrophylla (= V. ×subaffinis House).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 136. FNA vol. 6, p. 120.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. simulata, V. superba V. affinis var. subarctica, V. crenulata, V. sororia subsp. affinis, V. sororia var. affinis, V. subviscosa, V. venustula
Name authority Fischer ex Gingins: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 1: 296. (1824) Leconte: Ann. Lyceum Nat. Hist. New York 2: 138. (1826)
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