Viola labradorica
Viola vallicola
alpine violet, American dog or Labrador or alpine violet, American dog violet, dog violet, Labrador violet, violette du Labrador
sagebrush violet, valley violet, valley yellow violet, yellow prairie violet, yellow sagebrush violet
1–5, erect (later reclining), glabrous, on caudex from subligneous rhizome.
1–5, decumbent or ascending to erect, leafy proximally and distally, ca. 1/2 subterranean, glabrate to puberulent, on caudex from usually vertical, subligneous rhizome.
basal and cauline;
basal: 1–5;
stipules mostly lanceolate-linear, margins usually laciniate with gland-tipped projections, apex acute;
petiole 1–9.5 cm, glabrous or puberulent;
blade reniform or ovate to deltate, 1.4–5.5 × 1.8–5.3 cm, base cordate, not decurrent on petiole, margins crenate or serrate, mostly eciliate, apex acute to obtuse, surfaces usually glabrous, sometimes pubescent adaxially near margin;
cauline similar to basal except: stipules lanceolate to ovate, margins ± entire or laciniate, projections often gland-tipped, apex acuminate;
petiole 0.5–3.6 cm, glabrous;
blade narrowly ovate to orbiculate, 1.8–2.9 × 1.5–3.1 cm, base deeply cordate to truncate, margins regularly crenate, apex obtuse to broadly rounded or apiculate.
basal and cauline;
basal: 1–6;
stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, sometimes divided into filiform processes;
petiole 3–10 cm, glabrous or puberulent;
blade ovate to oblong-ovate, 1.5–4.3 × 0.9–1.1 cm, base usually truncate, sometimes attenuate, margins entire or serrulate, sometimes sinuate, ciliate (sometimes only on proximal 1/2 of leaf), apex acute to obtuse, surfaces glabrous or puberulent;
cauline similar to basal except: stipules linear to linear-lanceolate, margins ± entire, apex acute;
petiole 1.5–9.2 cm;
blade ovate or lanceolate to elliptic, 2.3–4 × 1–2 cm, length ca. 2.2 times width, base usually ± truncate to subcordate, sometimes attenuate on new leaves, margins sinuate, apex acute.
5–9 cm, glabrous or pubescent.
3–11.5 cm, glabrous or puberulent.
sepals lanceolate, margins ciliate or eciliate, auricles not enlarged in fruit, 1–1.5 mm;
petals usually lavender-violet to violet on both surfaces, rarely white, lower 3 white basally, darker violet-veined, lateral 2 sparsely bearded, lowest 9–16 mm, spur white to pinkish violet, elongated, 4–8 mm, tip usually straight, sometimes curved up;
style head beardless; cleistogamous flowers unknown.
sepals lanceolate, margins eciliate, auricles 0.5–1 mm;
petals deep lemon-yellow adaxially or on both surfaces, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 9–14 mm, spur yellow, gibbous, 0.8–1.2 mm;
style head bearded; cleistogamous flowers axillary.
ovoid to ellipsoid, 5–7 mm, glabrous.
spherical, ca. 5 mm, glabrous or finely puberulent.
beige to bronze, 1.5–2 mm.
tan, 2.1–2.2 mm, elaiosome extending beyond and covering funiculus.
= 20.
= 12.
Viola labradorica
Viola vallicola
H. E. Ballard (1994) made a compelling case for including Viola conspersa within V. labradorica, which is followed here. Ballard (1992) did not recognize V. labradorica as occurring in Colorado. Several western herbaria have collections identified as V. labradorica (ASC, CS, KHD, RM, RMBL; http://swbiodiversity.org/seinet/index.pho). Labels on some collections at KHD imply that V. labradorica is considered synonymous with V. adunca var. bellidifolia.
Viola labradorica reportedly hybridizes with V. adunca, V. rostrata (= V. ×malteana House), and V. striata (= V. ×eclipes H. E. Ballard), and less frequently with V. walteri var. appalachiensis and var. walteri.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
D. M. Fabijan et al. (1987) distinguished two varieties of Viola vallicola based on geographic location and type of leaf flavonoids: 1) var. major (Hooker) Fabijan, occurring west of the Continental Divide with leaf flavonoids primarily kaempferol derivatives; and 2) var. vallicola occurring east of the Continental Divide with leaf flavonoids all apigenin derivatives. Fabijan et al. reported that var. major (valley violet) occurs in sagebrush flats, prairie grasslands, open forests, juniper woodlands, 400–2800 m, in Alberta, Colorado, Idaho, Montana, Nevada, Oregon, Utah, and Washington.
M. S. Baker (1957) noted that there is only a tendency for leaves of Viola vallicola to be wide with truncate bases; on some plants only a few leaves are truncate. Images of type specimens at NY show truncate leaf bases for most basal and cauline leaves, with only some cauline leaves being attenuate. The key to V. vallicola in D. M. Fabijan et al. (1987) described the cauline leaf bases as truncate with some later cauline leaves becoming cuneate. G. Davidse (1976) stated that the V. vallicola plants he studied had cordate to truncate leaf bases; he made no distinction between basal and cauline leaves. Baker stated that V. vallicola may have given rise to V. nuttallii through a doubling of its chromosomes.
Because intermediate leaf forms are found in areas where Viola vallicola and V. nuttallii are sympatric, some question the specific status of V. vallicola. No hybrids involving V. nuttallii and V. vallicola are known (D. M. Fabijan et al. 1987).
V. Harms (pers. comm.) reported that Viola vallicola is frequent in southern Saskatchewan grasslands, usually occurring with V. nuttallii and appearing the more frequent of the two.
Observed pollinators of Viola vallicola in the intermountain region include flies in the genera Bombylius Linnaeus and Eristalis Latreille (G. Davidse 1976).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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