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alpine violet, American dog or Labrador or alpine violet, American dog violet, dog violet, Labrador violet, violette du Labrador

Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet

Habit Plants perennial, caulescent, not stoloniferous, 5–20 cm. Plants perennial, caulescent, not stoloniferous, 3–20 cm.
Stems

1–5, erect (later reclining), glabrous, on caudex from subligneous rhizome.

1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex.

Leaves

basal and cauline;

basal: 1–5;

stipules mostly lanceolate-linear, margins usually laciniate with gland-tipped projections, apex acute;

petiole 1–9.5 cm, glabrous or puberulent;

blade reniform or ovate to deltate, 1.4–5.5 × 1.8–5.3 cm, base cordate, not decurrent on petiole, margins crenate or serrate, mostly eciliate, apex acute to obtuse, surfaces usually glabrous, sometimes pubescent adaxially near margin;

cauline similar to basal except: stipules lanceolate to ovate, margins ± entire or laciniate, projections often gland-tipped, apex acuminate;

petiole 0.5–3.6 cm, glabrous;

blade narrowly ovate to orbiculate, 1.8–2.9 × 1.5–3.1 cm, base deeply cordate to truncate, margins regularly crenate, apex obtuse to broadly rounded or apiculate.

basal and cauline;

basal: 1–6, bipinnately compound, leaflets 3–5;

stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate;

petiole 5–6.8 cm, glabrous or puberulent;

blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;

cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute;

petiole 0.9–4 cm;

blade 1.1–4.1 × 1–3.6 cm.

Peduncles

5–9 cm, glabrous or pubescent.

2–12.5 cm, glabrous or puberulent.

Flowers

sepals lanceolate, margins ciliate or eciliate, auricles not enlarged in fruit, 1–1.5 mm;

petals usually lavender-violet to violet on both surfaces, rarely white, lower 3 white basally, darker violet-veined, lateral 2 sparsely bearded, lowest 9–16 mm, spur white to pinkish violet, elongated, 4–8 mm, tip usually straight, sometimes curved up;

style head beardless; cleistogamous flowers unknown.

sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;

petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm;

style head bearded; cleistogamous flowers absent.

Capsules

ovoid to ellipsoid, 5–7 mm, glabrous.

spherical to oblong, 5–12 mm, glabrous.

Seeds

beige to bronze, 1.5–2 mm.

light brown, 2.8–3.3 mm.

2n

= 20.

= 24, 48.

Viola labradorica

Viola douglasii

Phenology Flowering Apr–Aug. Flowering Feb–Jul.
Habitat Rich, mesic to wet woods, swamps, bogs Vernally moist grassy slopes and flats, often serpentine soil (except Oregon)
Elevation 50–3000 m (200–9800 ft) 20–2300 m (100–7500 ft)
Distribution
from FNA
AL; CO; CT; DC; DE; GA; IL; IN; KY; MA; MD; ME; MI; MN; NC; ND; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; MB; NB; NL; NS; NT; ON; PE; QC; SK; SPM; Greenland
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

H. E. Ballard (1994) made a compelling case for including Viola conspersa within V. labradorica, which is followed here. Ballard (1992) did not recognize V. labradorica as occurring in Colorado. Several western herbaria have collections identified as V. labradorica (ASC, CS, KHD, RM, RMBL; http://swbiodiversity.org/seinet/index.pho). Labels on some collections at KHD imply that V. labradorica is considered synonymous with V. adunca var. bellidifolia.

Viola labradorica reportedly hybridizes with V. adunca, V. rostrata (= V. ×malteana House), and V. striata (= V. ×eclipes H. E. Ballard), and less frequently with V. walteri var. appalachiensis and var. walteri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 135. FNA vol. 6, p. 129.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. adunca var. minor, V. canina var. muhlenbergii, V. conspersa, V. leucopetala, V. muhlenbergiana, V. muhlenbergiana var. minor, V. muhlenbergii V. chrysantha
Name authority Schrank: Denkschr. Kõnigl.-Baier. Bot. Ges. Regensburg 1(2): 12. (1818) Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841)
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