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Howell's violet, violette de Howell

cream or cream-white or pale or striped or striped cream violet, creamy violet, striped cream violet, striped white violet, violette strièe

Habit Plants perennial, caulescent, not stoloniferous, 2–44 cm. Plants perennial, caulescent, not stoloniferous, 10–60 cm.
Stems

1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome.

1–4, ascending to erect (often declining during anthesis), glabrous or pubescent, on caudex from fleshy rhizome.

Leaves

basal and cauline;

basal: 1–6;

stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate;

petiole 4–15 cm, glabrous or sparsely puberulent;

blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent;

cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate;

petiole 1–4.5 cm;

blade 2–5.1 × 1.2–5.8 cm.

basal and cauline;

basal: 2–6;

stipules lanceolate to narrowly deltate, margins laciniate, apex acute;

petiole 3–6 cm, glabrous or puberulent;

blade ovate to reniform, 2–7 × 1–2.5 cm, base cordate, margins crenate to serrate, ciliate or eciliate, apex obtuse to acute, surfaces glabrous or pubescent;

cauline similar to basal except: stipules lanceolate, margins laciniate;

petiole 3–7 cm;

distal blades ovate to deltate, 1–6 × 1–4 cm, base cordate, apex acuminate to acute.

Peduncles

2.8–17.8 cm, glabrous or sparsely pubescent.

5–12 cm, glabrous or pubescent.

Flowers

sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm;

petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight;

style head bearded; cleistogamous flowers axillary.

sepals lanceolate, margins ciliate or eciliate, auricles 2–3 mm;

petals white or cream on both surfaces, without yellow patch basally, lowest and usually lateral 2 purple-veined, lateral 2 and sometimes all densely bearded, lowest 10–18 mm, spur white, gibbous to elongated, 3–6 mm;

style head bearded; cleistogamous flowers axillary.

Capsules

ellipsoid, 7–11 mm, glabrous.

ellipsoid, 6–7 mm, glabrous.

Seeds

light brown, 2.1–2.8 mm.

beige to bronze, 1.5–3 mm.

2n

= 40, 80.

= 20.

Viola howellii

Viola striata

Phenology Flowering Apr–Jul. Flowering Mar–Jun.
Habitat Moist, shady areas, coniferous forests, prairies, along streams Riparian or alluvial woods, floodplains in silty loam, meadows
Elevation 50–1500 m (200–4900 ft) 40–1000 m (100–3300 ft)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KY; MA; MD; MI; MO; NC; NH; NJ; NY; OH; OK; PA; SC; TN; VA; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dead, long-persistent stems of Viola striata are often present the following season. In flower, the plants are often mistaken for V. canadensis because the petals are whitish (H. E. Ballard 1992).

Viola striata reportedly hybridizes with V. labradorica (= V. ×eclipes H. E. Ballard), V. rostrata (= V. ×brauniae Grover ex Cooperrider), V. walteri var. appalachiensis (= V. ×wujekii H. E. Ballard), and V. walteri var. walteri (= V. ×cooperrideri H. E. Ballard).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 134. FNA vol. 6, p. 159.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms Lophion striatum, V. conspersa var. masonii
Name authority A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887) Aiton: Hort. Kew. 3: 290. (1789)
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