Viola howellii |
Viola sheltonii |
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Howell's violet, violette de Howell |
cut-leaf violet, fan violet, Shelton's violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 2–44 cm. | Plants perennial, caulescent, not stoloniferous, 3–27 cm. |
Stems | 1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome. |
1–3, prostrate, decumbent, or erect, glabrous or sparsely puberulent, from short, often vertical, deep-seated or usually shallow, subligneous rhizome. |
Leaves | basal and cauline; basal: 1–6; stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate; petiole 4–15 cm, glabrous or sparsely puberulent; blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent; cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate; petiole 1–4.5 cm; blade 2–5.1 × 1.2–5.8 cm. |
basal and cauline; basal: 1–3, palmately compound, leaflets 3; stipules lanceolate-ovate, margins laciniate with gland-tipped projections, apex acute to acuminate; petiole 8.6–21 cm, glabrous or sparsely puberulent; blade reniform or ovate to ± orbiculate, 2–7 × 2–11 cm, coriaceous, base tapered, each leaflet cleft or dissected into 3 ± obovate lobes, each lobe further divided into 2–3 oblanceolate, pandurate, spatulate, oblong, lanceolate, or elliptic, lobes 2–10 mm wide, margins entire, ciliate or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or sparsely puberulent; cauline similar to basal except: stipules ovate to lanceolate, margin projections gland-tipped or eglandular, apex long-acuminate; petiole 5.5–12 cm; blade 1.2–6.3 × 1.2–10.5 cm. |
Peduncles | 2.8–17.8 cm, glabrous or sparsely pubescent. |
5–19 cm, glabrous or sparsely puberulent. |
Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm; petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins ciliate or eciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 dark brown to brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded or beardless, lowest 7–18 mm, spur yellowish with brownish purple specks, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary. |
Capsules | ellipsoid, 7–11 mm, glabrous. |
oblong to ovoid, 6–8 mm, glabrous or puberulent. |
Seeds | light brown, 2.1–2.8 mm. |
brownish, shiny, ca. 2.5 mm. |
2n | = 40, 80. |
= 12. |
Viola howellii |
Viola sheltonii |
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Phenology | Flowering Apr–Jul. | Flowering Mar–Jul. |
Habitat | Moist, shady areas, coniferous forests, prairies, along streams | Red fir, yellow pine, mixed evergreen, chaparral, oak woodlands, rich or gravelly soil |
Elevation | 50–1500 m (200–4900 ft) | 800–2500 m (2600–8200 ft) |
Distribution |
CA; OR; WA; BC
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CA; CO; ID; OR; WA
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Discussion | Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The cleistogamous flowers of Viola sheltonii are borne on long, prostrate peduncles usually buried in duff around the plant. Mature cleistogamous capsules are usually hidden and the dehisced seeds remain close to the parent plant. Some populations of V. sheltonii produce only cleistogamous flowers (D. Klaber 1976). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 134. | FNA vol. 6, p. 158. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. sheltonii var. biternata | |
Name authority | A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887) | Torrey: in War Department [U.S.], Pacif. Railr. Rep. 4(5): 67, plate 2. (1857) |
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