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Howell's violet, violette de Howell

goosefoot violet, goosefoot yellow violet, oakwoods violet

Habit Plants perennial, caulescent, not stoloniferous, 2–44 cm. Plants perennial, caulescent, not stoloniferous, 4–25(–34.5) cm.
Stems

1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome.

1–5, spreading to erect, leafy proximally and distally, usually elongated by end of season, puberulent to canescent, on caudex from subligneous rhizome.

Leaves

basal and cauline;

basal: 1–6;

stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate;

petiole 4–15 cm, glabrous or sparsely puberulent;

blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent;

cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate;

petiole 1–4.5 cm;

blade 2–5.1 × 1.2–5.8 cm.

basal and cauline;

basal: 1–6;

stipules adnate to petiole, forming 2 linear, membranous wings, each wing with lanceolate to ± deltate projection, margins entire or laciniate, apex usually long-acuminate or divided into narrow, filiform processes;

petiole 1.9–9.5 cm, puberulent;

blade usually grayish green to whitish, sometimes ± purple-tinted abaxially, green to grayish green adaxially, ± ovate to ± orbiculate, deltate, pandurate, or broadly brownish purple abaxially, lower 3 dark brown-veined, lateral 2 sparsely to densely bearded, lowest 10–16 mm, spur yellow to reddish brown, gibbous, 1–2 mm;

style head bearded; cleistogamous flowers axillary.

Peduncles

2.8–17.8 cm, glabrous or sparsely pubescent.

Flowers

sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm;

petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight;

style head bearded; cleistogamous flowers axillary.

Capsules

ellipsoid, 7–11 mm, glabrous.

ovoid to ellipsoid, 8–12 mm, puberulent.

Seeds

light brown, 2.1–2.8 mm.

medium brown, ± 2.7 mm.

2n

= 40, 80.

= 24.

Viola howellii

Viola quercetorum

Phenology Flowering Apr–Jul. Flowering Feb–Jul.
Habitat Moist, shady areas, coniferous forests, prairies, along streams Dry, grassy or brushy slopes, chaparral, in or lower than yellow pine forests
Elevation 50–1500 m (200–4900 ft) 300–2000 m (1000–6600 ft)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
Discussion

Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Herbarium specimens of Viola quercetorum and V. purpurea var. purpurea can be difficult to distinguish; pressing obliterates the undulate leaf margins of V. quercetorum. In Oregon, most collections of V. quercetorum are from yellow pine forests (A. Liston, pers. comm.).

Viola quercetorum hybridizes with V. douglasii (J. Clausen 1964).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 134. FNA vol. 6, p. 153.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. purpurea subsp. quercetorum
Name authority A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887) M. S. Baker & J. C. Clausen: Leafl. W. Bot. 5: 101. (1948)
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