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Howell's violet, violette de Howell

goosefoot violet, mountain violet, pine violet, purple-mark yellow violet

Habit Plants perennial, caulescent, not stoloniferous, 2–44 cm. Plants perennial, caulescent, not stoloniferous, 1.5–25 cm.
Stems

1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome.

1–5(–7), decumbent or spreading to erect, leafy proximally and distally, ± glabrous, puberulent, canescent, or tomentose, on caudex from subligneous rhizome.

Leaves

basal and cauline;

basal: 1–6;

stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate;

petiole 4–15 cm, glabrous or sparsely puberulent;

blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent;

cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate;

petiole 1–4.5 cm;

blade 2–5.1 × 1.2–5.8 cm.

basal and cauline;

basal: 1–6;

stipules adnate to petiole, forming 2 linear, membranous wings, wing margins entire or laciniate, each wing with lanceolate to ± deltate projection, margins entire or laciniate, apex acute to long-acuminate;

petiole 1.8–14.5 cm, puberulent to tomentose;

blade purplish, purple-tinted, or gray-green abaxially, gray, green, or gray-green adaxially, sometimes shiny adaxially, ovate, orbiculate, oblong, deltate, or lanceolate, 0.8–5.3 × 0.4–4.1 cm, often fleshy, base cordate, subcordate, truncate, or attenuate, oblique or not, margins usually ± crenate, serrate, dentate, or coarsely or irregularly repand-dentate, sometimes entire, usually ciliate, apex acute to obtuse or rounded, surfaces glabrous, puberulent, or tomentose;

cauline similar to basal except: stipules linear, lanceolate, oblanceolate, or ± oblong to ovate, margins entire, lacerate, or laciniate, usually ciliate, apex sometimes divided into 2–3 filiform processes or obtuse;

petiole 0.3–19.7 cm, glabrous or puberulent;

blade ovate, oblong, elliptic, deltate, or lanceolate, 0.9–5.2 × 0.2–2.9 cm, length 0.8–7.1 times width, margins crenate, serrate, dentate, repand-denticulate, undulate-denticulate, sinuate, undulate, or entire, abaxial surface puberulent, canescent, or tomentose, adaxial surface glabrous, sparsely pubescent, puberulent, canescent, or tomentose.

Peduncles

2.8–17.8 cm, glabrous or sparsely pubescent.

Flowers

sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm;

petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight;

style head bearded; cleistogamous flowers axillary.

Capsules

ellipsoid, 7–11 mm, glabrous.

ovoid to ± spherical, 4–7 mm, puberulent.

Seeds

light brown, 2.1–2.8 mm.

light to dark brown or mottled gray and brown, 2–3.1 mm.

2n

= 40, 80.

= 12.

Viola howellii

Viola purpurea

Phenology Flowering Apr–Jul.
Habitat Moist, shady areas, coniferous forests, prairies, along streams
Elevation 50–1500 m (200–4900 ft)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NV; OR; UT; WA; WY; BC; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 7 (7 in the flora).

Varieties of Viola purpurea are variable and intergrade. All are found in California; six occur in other western states, one in Mexico, and one in British Columbia. Mature plants are needed for determination.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal and cauline leaf blade surfaces tomentose
var. aurea
1. Basal and cauline leaf blade surfaces ± glabrous, canescent, or puberulent
→ 2
2. Stems mostly buried, not much elongated by end of season; plants 1.5–9(–12) cm
→ 3
2. Stems usually not buried, usually elongated by end of season; plants 3–25 cm
→ 4
3. Cauline leaf margins usually entire, sometimes sinuate; basal leaf margins ± crenate to irregularly repand-dentate or entire.
var. integrifolia
3. Cauline leaf margins usually coarsely crenate or dentate, sometimes ± serrate or ± entire; basal leaf margins coarse-serrate or irregularly dentate or crenate with 2–4 rounded teeth per side
var. venosa
4. Basal leaf base cordate or truncate
var. dimorpha
4. Basal leaf base usually attenuate, sometimes subcordate or truncate
→ 5
5. Cauline leaf blade length 3.2–7.1 times width, margins usually undulate-denticulate, sometimes entire
var. mesophyta
5. Cauline leaf blade length 1–3 times width, margins with 3–4(–5) pointed or rounded teeth per side or crenate-serrate
→ 6
6. Basal leaf blade margins irregularly crenate, without pointed or rounded teeth; adaxial surface often shiny.
var. purpurea
6. Basal leaf blade margins dentate-serrate with 4–5(–6) pointed or rounded teeth per side; adaxial surface not shiny.
var. mohavensis
Source FNA vol. 6, p. 134. FNA vol. 6, p. 150.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. purpurea var. aurea, V. purpurea var. dimorpha, V. purpurea var. integrifolia, V. purpurea var. mesophyta, V. purpurea var. mohavensis, V. purpurea var. purpurea, V. purpurea var. venosa
Name authority A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887) Kellogg: Pacific (San Francisco), 2 Feb. 1855: unnumb. (1855)
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