Viola howellii |
Viola praemorsa |
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Howell's violet, violette de Howell |
Astoria violet, Canary violet, marsh violet, upland yellow violet, yellow montane violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 2–44 cm. | Plants perennial, caulescent, not stoloniferous, 5.5–36.5 cm. | ||||||||
Stems | 1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome. |
1–3(–5), prostrate, decumbent, or erect, leafy proximally and distally, glabrous or puberulent, on caudex from usually vertical, subligneous rhizome. |
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Leaves | basal and cauline; basal: 1–6; stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate; petiole 4–15 cm, glabrous or sparsely puberulent; blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent; cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate; petiole 1–4.5 cm; blade 2–5.1 × 1.2–5.8 cm. |
basal and cauline; basal: 1–5; stipules adnate to proximal 1/3 of petiole, forming 2 narrow, linear-lanceolate wings, margins entire, apex of each wing free, acute; petiole 2.6–19.2 cm, glabrous or densely puberulent; blade usually elliptic to ovate, sometimes oblong-lanceolate to ± orbiculate, 1.7–14(–17) × 0.8–5.3(–6.7) cm, base attenuate to ± truncate or subcordate, often oblique, margins usually crenate, serrulate, or serrate, sometimes entire, ciliate or eciliate, apex acute to obtuse, surfaces glabrous or densely puberulent; cauline similar to basal except: stipules lanceolate to ovate, margins entire or toothed, with or without gland-tipped projections, apex acute to acuminate; petiole 1.3–16.2 cm, glabrous or puberulent; blade 2.3–11(–14.8) × 1.1–3.6(–5.5) cm, length 1.1–6.5 times width. |
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Peduncles | 2.8–17.8 cm, glabrous or sparsely pubescent. |
4.4–27 cm, glabrous or puberulent. |
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Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm; petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins eciliate, auricles 1–2 mm; petals deep lemon-yellow adaxially, upper 2, and sometimes lateral 2, brownish purple abaxially, lower 3 brownish purple-veined, lateral 2 sparsely bearded, lowest 11–19 mm, spur yellow or pale green, gibbous, 0.5–3 mm; style head bearded; cleistogamous flowers axillary. |
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Capsules | ellipsoid, 7–11 mm, glabrous. |
ellipsoid to oblong, 6–14 mm, glabrous or finely puberulent. |
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Seeds | light brown, 2.1–2.8 mm. |
medium to dark brown or red-brown, 2–3 mm, elaiosome completely covering funiculus. |
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2n | = 40, 80. |
= 36, 48. |
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Viola howellii |
Viola praemorsa |
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Phenology | Flowering Apr–Jul. | |||||||||
Habitat | Moist, shady areas, coniferous forests, prairies, along streams | |||||||||
Elevation | 50–1500 m (200–4900 ft) | |||||||||
Distribution |
CA; OR; WA; BC
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CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
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Discussion | Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). Viola praemorsa is a complex group that appears to be evolving. Sympatric populations and the similarity of their flowers provide opportunities for hybridization and introgression (D. M. Fabijan et al. 1987). Some botanists have suggested that all taxa in the V. nuttallii complex, which includes V. bakeri, V. nuttallii, V. praemorsa, V. tomentosa, and V. vallicola, should be treated as varieties of V. nuttallii, as C. L. Hitchcock et al. (1955–1969, vol. 3) did, on the basis that these taxa are more or less sympatric in range, intergrade with one another, and possess no distinctive gross morphological features by which they can be consistently recognized. S. L. Welsh et al. (1987) treated V. praemorsa and V. linguaefolia Nuttall as synonymous with V. nuttallii, stating that V. nuttallii (and related taxa) are acaulescent to short-caulescent. Although some plants have short internodes, all plants in the V. nuttallii complex are caulescent. We acknowledge the complexity and taxonomic difficulties inherent in the V. nuttallii complex; patterns of morphology, cytology, and leaf flavonoid chemistry provide a basis for recognizing infrataxa. Fabijan et al. conducted the most extensive study of the complex to date; their treatment of V. praemorsa is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 134. | FNA vol. 6, p. 145. | ||||||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||||||
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Synonyms | V. nuttallii subsp. praemorsa, V. nuttallii var. praemorsa | |||||||||
Name authority | A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887) | Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1254. (1829) | ||||||||
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