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Howell's violet, violette de Howell

pinto violet, two-eyed violet, western heart's ease, western heart's ease or two-eyed or southern two-eyed violet

Habit Plants perennial, caulescent, not stoloniferous, 2–44 cm. Plants perennial, caulescent, not stoloniferous, 2–37 cm.
Stems

1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome.

1–3, ascending to erect, ± glabrous or usually puberulent, on caudex from shallow, fleshy rhizome or deep-seated caudex with fleshy roots.

Leaves

basal and cauline;

basal: 1–6;

stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate;

petiole 4–15 cm, glabrous or sparsely puberulent;

blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent;

cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate;

petiole 1–4.5 cm;

blade 2–5.1 × 1.2–5.8 cm.

basal and cauline;

basal: 1–6;

stipules deltate, margins laciniate, apex usually long-acuminate;

petiole 3.7–10 cm, puberulent;

blade ovate, deltate, or subreniform, 1–6 × 1.2–4 cm, base usually cordate to subcordate, margins crenate, usually ciliate, apex acute, mucronulate, surfaces usually sparsely puberulent, glabrous early;

cauline similar to basal except: stipules lanceolate, margins ±fimbriate, sometimes stipitate-glandular, sometimes entire, erose, or ± laciniate;

petiole 0.4–9 cm;

blade ovate to deltate, 1.6–4.4 × 1.1–3.6 cm, base subcordate to truncate, margins crenate to ± serrate, ciliate, apex acute to obtuse.

Peduncles

2.8–17.8 cm, glabrous or sparsely pubescent.

1–10 cm, puberulent.

Flowers

sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm;

petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight;

style head bearded; cleistogamous flowers axillary.

sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;

petals white adaxially, upper 2 and sometimes lower 3 deep reddish violet abaxially, all with yellow area basally, lateral 2 with purple patch basally distal to smaller yellow area, bearded, lowest with yellow patch basally, purple-veined, 5–15 mm, spur yellow or greenish, gibbous, 1–2.5 mm;

style head bearded; cleistogamous flowers axillary.

Capsules

ellipsoid, 7–11 mm, glabrous.

spherical to spherical-ovoid, 5–11 mm, minutely scabrous.

Seeds

light brown, 2.1–2.8 mm.

brown-purple, ca. 2 mm.

2n

= 40, 80.

= 12.

Viola howellii

Viola ocellata

Phenology Flowering Apr–Jul. Flowering Mar–Jul.
Habitat Moist, shady areas, coniferous forests, prairies, along streams Rocky areas, grassy banks, thickets, often on serpentine soil
Elevation 50–1500 m (200–4900 ft) 100–1100 m (300–3600 ft)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 134. FNA vol. 6, p. 140.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Name authority A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887) Torrey & A. Gray: Fl. N. Amer. 1: 142. (1838)
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